Silene neglecta Ten., Fl. Neapol. Prodr. App. 5: 13. 1826.

Saez, Llorenc, Mesbah, Melilia, Lopez-Alvarado, Javier, Bacchetta, Gianluigi, Mokni, Ridha El, Peruzzi, Lorenzo & Oxelman, Bengt, 2022, Re-establishment of Silene neglecta Ten. (Caryophyllaceae) with taxonomic notes on some related taxa, PhytoKeys 195, pp. 143-160 : 143

Silene neglecta Ten., Fl. Neapol. Prodr. App. 5: 13. 1826.

Silene nocturna neglecta ≡ Silene nocturna subsp. neglecta (Ten.) Arcang. Comp. Fl. Ital.: 88. 1882. Lectotype (designated by Peruzzi & Carta, 2013: 45): [Italy] Campania: Persano, s.d., Tenore s.n. (NAP!).

Silene martinolii = Silene martinolii Bocchieri & B.Mulas, Boll. Soc. Sarda Sci. Nat. 26: 301. 1988; syn. nov. Holotype (Bocchieri & Mulas, 1988: 301): [Italy. Sardinia] Isola il Toro, 22 May 1988, Bocchieri & Mulas s.n. (CAG!).

Description.

Annual herb (rarely biennial) green or greyish-green, hairy. Stems (10-)15-50(-70) cm, more or less erect (rarely prostrate-ascending), unbranched to much-branched, densely hairy, usually viscid above. Lower leaves spatulate to obovate; cauline leaves obovate to linear-lanceolate. Flowers 2-9(-10) in raceme-like monochasial cymes; lowermost pedicels shorter or longer than calyx (0.45-2.00 times as long as calyx in fruit), erect to patent, pubescent-glandular. Calyx 8-13 mm long, subcylindrical to cylindrical-ovoid in flower, attenuate, becoming subovoid to ovoid in fruit, usually densely hairy, covered by long eglandular hairs (up to 1.9 mm long) and glandular hairs (up to 2 mm long); calyx teeth triangular to linear-lanceolate or linear, acute; veins anastomosing. Petals pale pink to pinkish-purple, rarely white; limb 4.5-6.0 mm, subentire or emarginate; coronal scales 1.2-2.2 mm, whitish to pink. Stamens with alternate filaments hairy at base, sometimes all hairy at base; anthers 1.0-1.5 mm long, lilac to purple, exserted from corolla mouth. Gonophore (1.8-)2.0-2.4(-2.6) mm long, puberulent. Capsule 6.0-8.5 mm, subcylindrical, enclosed within the calyx. Seeds (0.90-)0.91-1.02(-1.05) × (0.76-)0.80-0.89(-0.91) blackish to dark brown; faces deeply concave, tuberculate; back wide, slightly canaliculate.

Flowering time.

Between March (exceptionally at the beginning of February) and May.

Chromosome number.

2 n = 24 ( Peruzzi and Carta 2013; Bosch et al. 2019).

Habitat.

Rocky places, maritime sands and grassland, usually on siliceous substrata, 0-1700 m a.s.l.

Distribution.

Central and southern Italy, south-western Sardinia, Sicily, northern Algeria and Tunisia and north-eastern Spain.

Specimens examined.

Algeria: Terrains siliceux, berges maritimes sur la route de Cap de Garde, au-dessous de la colline des Caroubiers à Bône, 5 March 1867, Tribout (P05032571); Bône, rochers maritimes, 12 Apr 1867, Tribout (P05033844); subéraire, près de Bessombourg, 500-600 m alt., Kabylie de Collo, 17 May 1944, L. Faurel (P05110031, P05135518); subéraire, des crètes de Boudra, près de Bessombourg, 600 m alt., Kabylie de Collo, 26 May 1944, L. Faurel (P05050465); pentes rocheuses du flanc sud du Djebel Tababort, 1700 m alt., (Dt. de Constantine), 29 May 1952, L. Faurel (P05110033); près du Col de Terras, 800 m alt., environ 5 km à l’ouest de Zitouna (ex Bessombourg), Kabylie de Collo, 8 Juin 1984, A. Dubuis (MPU286766); Italy: Campania: Ischia, Oct 1835, Gussone (K00728586); Campania, vigneti del Vesuvio, 150 m alt., 29 Apr 1911, G. Pellanda (BC8940, MA 31093, US1272504); Campi Flegrei, prope Capo Miseno, 100 m alt., solo siliceo, 15 May 1913, M. Guadagno (BC8939); Torre Gaveta, May 1913, M. Guadagno (MA31092, MA31096, MPU300592); Sardinia: Isola il Toro, 22 May 1988, E. Bocchieri & B. Mulas (CAG, holotype S. martinolii ); Isola Rossa, Golfo di Teulada, 10 July 1988, E. Bocchieri (CAG, sub S. martinolii ); Isola la Vacca, 18 Feb 1990, E. Bocchieri (CAG, sub S. martinolii ); S’Aqua sa Canna, Isola di S. Antioco, 12 Apr 1992, L. Mossa (CAG, sub S. martinolii ); Sa Corona su Crabi, isola di S. Antioco, 20 May 1993, L. Mossa (CAG, sub S. martinolii ); Sicily: Lipari, Apr 1902, Ross 317 (WAG); Vulcano Island, Apr 2017, D. Carrera (grown from seed by J. López & L. Sáez, May 2018, L. Sáez, herb. pers.); Tuscany: Insula Igilium (Giglio), 17 May 1894 Sommier (MPU772254); Insula Igilium (hod. Giglio), prope Portum, 5 Apr 1899, Sommier (MA31097); Insula Inarime, in campis aridis, May 1905, M. Guadagno ( US548489 ); Isola d’Elba (Tuscan Archipelago), Miniera del Ginevro, 27 Apr 2017, B. Pierini (PI, sub S. mutabilis ). Spain: Barcelona: Bruguers, 25 May 1984, C. Blanché ( BCN 41111 ); Garraf massif, Gavà, Ermita de Bruguers, siliceous rocks, 265 m alt., 6 May 2015, L. Sáez LS-7626 (L. Sáez, herb. pers.); Cervelló, Can Riera, 350 m alt., 9 May 2015, D. Carrera & L. Sáez LS-7629 (L. Sáez, herb. pers.); Torrelles de Llobregat, Roca Plana NE from Turó de la Bruguera, 280 m alt., 23 Apr 2016, L. Sáez LS-7706 (L. Sáez, herb. pers.); Gavà, Roca del Barret, 23 Apr 2016, L. Sáez (L. Sáez, herb. pers.); ibidem, 26 May 2016, L. Sáez LS-7707 (L. Sáez, herb. pers.); ibidem, 29 Apr 2021, L. Sáez (herb. pers.); between castell d’Eramprunyà and Bruguers, 26 May 2016, L. Sáez (herb. pers.). Tunisia: Tabarka, maritime sands, March 2017, R. El Mokni (L. Sáez, herb. pers.; R. El Mokni, herb. pers.).