Megasoma (Lycophontes) joergenseni penyai Nagai, 2003

Prandi, Massimo, Grossi, Paschoal C. & Vaz-De-Mello, Fernando Z., 2020, Revision of the Megasoma (Megasoma) gyas (Jablonsky in Herbst, 1785) species group (Coleoptera, Scarabaeidae, Dynastinae), ZooKeys 999, pp. 109-145 : 109

Megasoma (Lycophontes) joergenseni penyai Nagai, 2003 Figure 27

Megasoma joergenseni ssp. penyai Nagai, 2003: 38-39.

Remarks.

The subspecies penyai is restricted to an area in central-western Paraguay (holotype from Loma Plata, Chaco). The main characters differentiating it from the subspecies joergenseni are found in the smaller median thoracic horn and in the denser pubescence, giving a more brownish color. It is also usually smaller than ssp. joergenseni . (Fig. 27 , courtesy of Mushi-sha).

Figures

Figure 27. A M. joergenseni peynai ♂ B M. joergenseni peynai ♀.