Harpalus Latreille, 1802

Bousquet, Yves, 2012, Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico, ZooKeys 245, pp. 1-1722 : 699

Subgenus Harpalus Latreille, 1802

Harpalus Latreille, 1802: 92. Type species: Carabus proteus Paykull, 1790 (= Carabus affinis Schrank, 1781) designated by Andrewes (1935: 19).

Proteonus Fischer von Waldheim, 1829a: 21. Type species: Carabus distinguendus Duftschmid, 1812 designated by Bousquet (2002c: 178). Synonymy established by Bousquet (2002c: 178).

Amblystus Motschulsky, 1864: 209. Type species: Carabus rubripes Duftschmid, 1812 by original designation. Synonymy established by Tschitschérine (1901: 240).

Harpalomerus Casey, 1914: 76. Type species: Harpalus amputatus Say, 1830 designated by Lindroth (1968: 769). Synonymy established by Kataev et al. (2003: 371). Etymology. From the generic name Harpalus [q.v.] and the Greek meros (part, segment) [masculine].

Diversity.

This subgenus includes native species in North America, Mexico, and the Palaearctic and Afrotropical Regions. The total number of species cannot be determined at this time. The North American fauna is represented by 17 species, of which two are adventive.

Identification.

Noonan (1991) revised all the North American species except the two adventive ones but his key (pages 20-45) included all 17 species. Lindroth (1968) covered all but three ( Harpalus balli , Harpalus martini , and Harpalus rubripes ) species in his monograph of the Canadian Carabidae .