Cratocerus kavanaughi Grzymala & Will

Grzymala, Traci L. & Will, Kipling W., 2014, Taxonomic review of Cratocerus Dejean, 1829 (Coleoptera, Carabidae) with the description of six new species, ZooKeys 416, pp. 77-112 : 94-95

Taxon classification Animalia Coleoptera Carabidae

Cratocerus kavanaughi Grzymala & Will sp. n. Figs 7, 9g, 10e, 11f, 12c

Type locality.

French Guiana, Nouragues Scientific Station, Pararé site, coordinates 4.02N, 52.41W.

Type material.

HOLOTYPE [male, deposited in NMNH] labeled: "FRENCH GUYANA: Nouragues - / Saut Parare, Commune de Regina / 4.02°N, 52.41°W, 51 meters / 26 Apr. 2010, flight intercept trap / S. Brule, P.H. Dalens & E. Poirier // ADP / 126766 // U.C. Berkeley / EMEC / 654851".

Paratypes. 10 total; 1 female, labeled: "FRENCH GUYANA: Nouragues - Saut Parare, Commune de Regina 4.02°N, 52.41°W, 51 meters 20 Apr. 2010, flight intercept trap S. Brule, P.H. Dalens & E. Poirier (EMEC207921); 1 female, labeled: "FRENCH GUYANA: Nouragues - Saut Parare, Commune de Regina 4.0378°N, 057.6725°W, 51m. 30 Nov. 2009, flight intercept trap S. Brule, P.H. Dalens & E. Poirier" (EMEC207901); 1 male, labeled: "FRENCH GUYANA: Reserve Naturelle des Nouragues Inselberg, Camp, Commune de Regina, 4°05'N, 52°41'W, flight intercept trap 23 February 2011 S. Brule, P.H. Dalens & E. Poirier" (EMEC654857); 1 male, labeled: "FRENCH GUYANA: Montagne des Chevaux Commune de Roura, RN2 PK22, 4.7127°N, 52.3966°W, 90 meters 21 Dec. 2011, flight intercept trap S. Brule, P.H. Dalens & E. Poirier" (EMEC654907); 1 female, labeled: "FRENCH GUYANA: Montagne des Chevaux Commune de Roura, RN2 PK22, 4.7127°N, 52.3966°W, 900 m. 26 Dec. 2010, flight intercept trap S. Brule, P.H. Dalens & E. Poirier" (EMEC654852); 1 female, labeled: "FRENCH GUYANA: Montagne des Chevaux Commune de Roura, RN2 PK22, 4.7127°N, 52.3966°W, 90 meters 23 July 2011, SLAM Malaise trap S. Brule, P.H. Dalens & E. Poirier" (EMEC654856); 1 female, labeled: "FRENCH GUYANA: Region de Saul, Commune de Saul, Belvedere de Saul, 3.6223N, 53.2159W, 283-325 meters 24 Jan. 2011, flight intercept trap, VPV S. Brule, P.H. Dalens & E. Poirier" (EMEC654853); 1 female, labeled: "FRENCH GUYANA: Region de Saul, Commune de Saul, Belvedere de Saul 3.6223N, 53.2159W, 283-325 meters 07 Apr. 2011, flight intercept trap, V6 S. Brule, P.H. Dalens & E. Poirier" (EMEC654854); 1 male, labeled: "FRENCH GUYANA: Region de Saul, Commune de Saul, Belvedere de Saul 3.6223N, 53.2159W, 283-325 meters 20 Feb.. 2010, flight intercept trap, V5 S. Brule, P.H. Dalens & E. Poirier" (EMEC654845); 1 male, labeled: "PERU: MADRE DE DIOS Pakitza, Zone 2 07Oct90 T.L. Erwin & M.G. Pogue Colls, 12°07'S, 70°58'W, Under bark on Tr. Aguajal nr. St. 127" (EMEC207851). All specimens deposited in NMNH except EMEC654857 deposited in MNHN.

Diagnosis.

This species is distinguished from other members of Cratocerus by the strongly curved mandibles, 45 degrees in relation to the mandibular midline and the two basal pronotal fovea. This species is most easily confused with Cratocerus culpepperi as both have the distinctively curved mandibles, a character which does not occur in any other species within Cratocerus . Cratocerus kavanaughi is easily identified by the larger overall body size and the two pronotal basal fovea as opposed to the five found with Cratocerus culpepperi specimens (Fig. 9g, h).

Description.

Size average for the genus, ABL 4.8-5.6 mm, TW 1.9-2.4 mm. Color. Head and prothorax brunneopiceous; elytra, abdominal sternites, and mandibles brunneous; legs, antennae, and remaining mouthparts rufotestaceous. Head. HL 0.7-1.0 mm, HW 0.8-0.9 mm. One supraorbital seta present over each eye. Apices of mandibles strongly curved, apices at more than 45 degree angle to mandibular midline. Apical maxillary palpi length 4 × width at base. Mentum with triangulate lateral lobes. Thorax. Pronotum (Fig. 9g) with distinct median impression, one distinct basal fovea laterad; one lateral seta located on apicolateral pronotal bead; one seta present on posterior bead and projected over posterior angle; pronotal anterior margin convex, greatest width anterad of center; pronotal width 1.3-1.4 × pronotal length. Elytral length 1.4-1.6 × elytral width. Legs. Protibia with three small, stout spines confined to distal half of lateral margin, only visible with extreme magnification (Fig. 10e). Genitalia, male. Median lobe curved sharply, between forty-five and ninety degrees with respect to basal 1/3, remaining 2/3 without continuing curvature; narrowed apically; length average for species of the genus. Left paramere curved and simple (Figure 11f). Genitalia, female. Spermatheca with entire length equal and moderate, length average for genus; apex rounded, without broad bulb (Fig. 12c).

Etymology.

The specific epithet, kavanaughi, is a Latinized version (genitive case) of the surname of Dr. David H. Kavanaugh, an entomology curator at the California Academy of Sciences. Dr. Kavanaugh provided numerous resources and support to TLG throughout the duration of this project. This species is named in gratitude of this support.

Distribution.

This species is geographically distributed (Fig. 13) throughout French Guiana and Peru.

Ecology.

Specimens have been collected in flight intercept traps during the months of November, December, January, February, April, and July and under tree bark in October.