Notocupes Ponomarenko, 1964

STRELNIKOVA, OLESYA D. & YAN, EVGENY V., 2023, Redescriptions of the Triassic Notocupes beetles (Archostemata: Ommatidae) from Kyrgyzstan and South Kazakhstan, Palaeoentomology 6 (2), pp. 174-190 : 176-177

Genus Notocupes Ponomarenko, 1964

Notocupes Ponomarenko, 1964: 61 , fig. 5.

Sinocupes Lin, 1976: 113 (type species: Sinocupes validus Lin, 1976 ); Ponomarenko, 2006: 90; Kirejtshuk et al., 2010: 792.

Chengdecupes Hong, 1983: 83 , fig. 75 (type species: Chengdecupes jurassicus Hong, 1983 ); Kirejtshuk et al., 2016: 144.

Conexicoxa Lin, 1986: 76 (type species: Conexicoxa homorus Lin, 1986 ); Ponomarenko, 2006: 90.

Forticupes Hong & Wang, 1990: 105 (type species: Forticupes laiyangensis Hong, 1990 ); Ponomarenko, 2006: 90.

Picticupes Hong & Wang, 1990: 107 (type species: Picticupes tuanwangensis Hong, 1990 ); Ponomarenko, 2006: 90.

Lupicupes Ren et al., 1995: 74 (type species: Lupicupes trachylenus Ren, 1995 ); Ponomarenko, 2006: 90.

Amblomma Tan et al., 2005: 208 (type species: Amblomma psilata Tan, Ren & Liu, 2005 ); Kirejtshuk et al., 2010: 788.

Ovatocupes Tan & Ren, 2006: 225 (type species: Ovatocupes alienus Tan & Ren, 2006 ); Kirejtshuk et al., 2010: 788.

Type species. Notocupes picturatus Ponomarenko, 1964 ; Upper Jurassic , Karabastau Formation , Karatau, Southern Kazakhstan .

Species included. The genus consists of 64 species. Among them species of synonymized genera: Ovatocupes Tan & Ren, 2006 , Sinocupes Lin, 1976 , Forticupes Hong & Wang, 1990 , Amblomma Tan et al., 2005 , Picticupes Hong & Wang, 1990 , Lupicupes Ren, 1995 , Conexicoxa Lin, 1986 , Chengdecupes Hong, 1983 and species transferred to Notocupes from Procarabus Oppenheim, 1888 and Tetraphalerus Waterhouse, 1901 .

Notocupes reticulatus ( Oppenheim, 1888) (= Procarabus reticulatus Oppenheim,1888 ), N. tripartitus ( Oppenheim, 1888) (= Procarabus tripartitus Oppenheim, 1888 ), N. picturatus Ponomarenko, 1964 , N. excellens Ponomarenko, 1966 , N. caudatus Ponomarenko, 1966 , N. vitimensis Ponomarenko, 1966 , N. reticulatus Ponomarenko, 1968 , N. lapidarius Ponomarenko, 1968 , N. nigrimonticola Ponomarenko, 1968 , N. pulcher Ponomarenko, 1968 , N. caducus Ponomarenko, 1969 , N. crassus Ponomarenko, 1969 , N. oxypygus Ponomarenko, 1969 , N. laticella Ponomarenko, 1969 , N. tenuis Ponomarenko, 1969 , N. rostratus Ponomarenko, 1969 , N. cellulosus Ponomarenko, 1969 , N. issykkulensis Ponomarenko, 1969 , N. latus Ponomarenko, 1969 , N. kirghizicus Ponomarenko, 1969 , N. sogutensis Ponomarenko, 1969 , N. laetus Lin, 1976 , N. validus ( Lin, 1976) (= Sinocupes validus Lin, 1976 ), N. jurassicus ( Hong, 1983) (= Chengdecupes jurassicus Hong, 1983 ), N. shiluoensis (Hong, 1984) (= Ch. shiluoensis Hong, 1984 ), N. patulus Ponomarenko, 1985 , N. dischides Zhang, 1986 , N. homorus ( Lin, 1986) (= Conexicoxa homorus Lin, 1986 ), N. kezuoensis (Hong, 1987) (= Ch. kezuoensis Hong, 1987 ), N. tuanwangensis ( Hong & Wang, 1990) (= Picticupes tuanwangensis Hong & Wang, 1990 ), N. laiyangensis ( Hong & Wang, 1990) (= Forticupes laiyangensis Hong & Wang, 1990 ), N. baojiatunensis (Hong, 1992) (= Ch. baojiatunensis Hong, 1992 ), N. khetanensis Ponomarenko, 1993 , N. brachycephalus Ponomarenko, 1994 , N. exiguus Ponomarenko, 1994 , N. longicollis Ponomarenko, 1994 , N. mongolicus Ponomarenko, 1994 , N. elegans Ponomarenko, 1994 , N. dundulaensis Ponomarenko, 1994 , N. trachylenus ( Ren, 1995) (= Lupicupes trachylenus Ren, 1995 ), N. lentus ( Ren, 1995) (= Tetraphalerus lentus Ren, 1995 ), N. ludongensis Wang & Liu, 1996 , N. semen ( Ponomarenko, 2000) (= Zygadenia semen Ponomarenko, 2000 ), N. sibirica ( Ponomarenko, 2000) (= Z. sibirica Ponomarenko, 2000 ), N. psilata ( Tan, Ren & Liu, 2005) (= Amblomma psilata Tan, Ren & Liu, 2005 ), N. rudis ( Tan, Ren & Liu, 2005) (= A. rudis Tan, Ren & Liu, 2005 ), N. stabilis ( Tan, Ren & Liu, 2005) (= A. stabilis Tan, Ren & Liu, 2005 ), N. epicharis ( Tan, Ren & Liu, 2005) (= A. epicharis Tan, Ren & Liu, 2005 ), N. alienus (Tan & Ren, 2006) (= Ovatocupes alienus Tan & Ren, 2006 ), N. viridis ( Soriano & Delclòs, 2006) (= Z. viridis Soriano & Delclòs, 2006 ), N. oculata ( Soriano & Delclòs, 2006) (= Z. oculata Soriano & Delclòs, 2006 ), N. diazromerali ( Soriano & Delclòs, 2006) (= Z. diazromerali Soriano & Delclòs, 2006 ), N. martinclosas ( Soriano & Delclòs, 2006) (= Z. martinclosas Soriano & Delclòs, 2006 ), N. longicoxa ( Soriano & Delclòs, 2006) (= Z. longicoxa Soriano & Delclòs, 2006 ), N. siniestri ( Soriano & Delclòs, 2006) (= Z. siniestri Soriano & Delclòs, 2006 ), N. cyclodonta (Tan et al., 2006) (= A. cyclodonta Tan et al., 2006 ), N. miniscula (Tan et al., 2006) (= A. miniscula Tan et al., 2006 ), N. porrecta (Tan et al., 2006) (= A. porrecta Tan et al., 2006 ), N. eumeura (Tan et al., 2006) (= A. eumeura Tan et al., 2006 ), N. protensa (Tan et al., 2006) (= A. protensa Tan et al., 2006 ), N. pingi Ponomarenko & Ren, 2010 , N. lini Ponomarenko et al., 2012 , N. khasurtyiensis Strelnikova, 2019 , and N. premeris Lee et al., 2022 .

Diagnosis. Beetle of medium size. Scapus notably larger than other ten segments; pedicellus usually equal in size to fourth or fifth one; rest of antennomeres gradually diminishing in size to the top. Pronotum transverse, its sides explanate, anterior angles at least slightly protruding anteriorly. Elytron with 10 rows of cells, organized in twos (except for cell rows on epipleuron). Two veins closest to the sutural margin (Cu and A2 sensu Jarzembowski et al., 2015 and 4 and 5 on Figs 2 С, 6D, 8D, 12) merging near elytral apex as short veins. Apical abdominal sternite 2– 4× as long as penultimate one; abdominal sternites I–IV overlapping each other.

Remarks. Four species of Pareuryomma Tan et al., 2012 genus, which was synonymized with Notocupes ( Kirejtshuk et al., 2010) , were later excluded as synonyms of Notocupes ( Tan et al., 2012; Kirejtshuk, 2020).

Ten species of Zygadenia Handlirsch, 1906 are excluded as synonyms of Notocupes as well, namely, Z. tuberculata ( Giebel,1856) (= Curculionitestuberculatus Giebel, 1856), Z. angliae ( Giebel, 1856) (= Kakoselia angliae Handlirsch, 1906 ), Z. westraliensis (Riek, 1968) (= Mesothoris westraliensis Riek, 1968 ), Z. sinitzae Ponomarenko, 2000 , Z. handlirschi Ponomarenko, 2014 , Z. giebeli Ponomarenko, 2014 , Z. liui Jarzembowski, 2015 , Z. floodpagei Jarzembowski, 2015 , Z. simpsoni Jarzembowski, 2015 , and Z. martinae Jarzembowski, 2015 . Zygadenia is treated here as a formal taxon for isolated elytra similar to Notocupes that may, or may not, belong to Notocupes .

Figures

FIGURE 2. Notocupes laticella, paratypes (PIN, 2240/71, 74). A, Wetted with 95% ethanol PIN, 2240/74. B, Interpretative line drawing (square here and beyond shows relative size and density of tubercles). С, General scheme of elytra. D, Wetted with 95% ethanol PIN, 2240/71, counterpart. E, Interpretative line drawing. F, Interpretative line drawing of PIN, 2240/71, part. G, Photo of dry PIN, 2240/71, part. Abbreviations here and elsewhere: in.cx.p, intercoxal process; mesan, mesanepisternum; mesep, mesepimeron, mesotr, mesotrochantin; mesovent, mesoventrite; metanep, metanepimeron; metatr, metatrochantin; metavent, metaventrite; mnd, mandible; P2, supraocular protuberance; P3, posteromesal protuberance; s.n.pl, notopleural suture; s.pl.st, pleurosternal suture.