Cratocerus tanyae Grzymala & Will

Grzymala, Traci L. & Will, Kipling W., 2014, Taxonomic review of Cratocerus Dejean, 1829 (Coleoptera, Carabidae) with the description of six new species, ZooKeys 416, pp. 77-112 : 89-92

Taxon classification Animalia Coleoptera Carabidae

Cratocerus tanyae Grzymala & Will sp. n. Figs 5, 9d, 10c, 11d

Type locality.

Costa Rica, Limon Province, Reserva Biológica Hitoy-Cerere, Sendero Bobocara. Estimated coordinates 9.67N, 83.03W.

Type material.

HOLOTYPE [male, deposited in INBio] labeled: "COSTA RICA. Prov. Limon Reserva / Biol Hitoy Cerere, Send Bobocara. / 200m. 13 ABR 2000. W. Arana / Mantillo L_N_184700_641200 #56335 // INB0003076608 / INBIOCRI COSTA RICA // U.C. Berkeley / EMEC 207,842".

Paratypes. 19 total; 1 female, labeled: "Sector Cerro Cocori, Finca de E. Rojas, Prov. Limon, COSTA RICA, 150m. SET 1994, E. Rojas, L_N_286000_567500 #3218" (EMEC207845); 1 female, labeled: "COSTA RICA. Prov. Limon, R.B. Hitoy Cerere, A.C.L.A.C. Estac. Hitoy Cerere, Send, Catarata, 200m, 30 SEP 2000. W. Arana, Mantillo con Sopa L_N_184550_643250 #63428" (EMEC207999); 1 male and 1 female, labeled: "Est. Quebrada Bonita, Prov. Punta, COSTA RICA. 50m. Nov 1994 R. Guzman, L N 194500_469850 #3258" (EMEC207990, EMEC207994); 1 male, labeled: "Est. Q. Bonita, Prov. Punta, COSTA RICA. 50m. Jun 1993. R. Guzman, L N 194500_469850 #2202" (EMEC207991); 1 male and 1 female, labeled: "Est. Queb. Bonita, 50m, Res. Biol. Carara, Prov. Puntarenas, Costa Rica Dic 1992, R. Guzman L_N 194500, 469850" (EMEC207992, EMEC207993); 1 male, labeled: "COS TA RICA. Prov. Limon, Est. R.B. Hitoy Cerere, La poza de las chichis, 140m. 18-29 JUN 1998. E. Rojas, Tp. Mantillo L_N_184176_643226 #50751" (EMEC207841); 1 female, labeled: "COSTA RICA. Prov. Puntarenas, P.N. Corcovado, Est. Sirena, Send, Espavel, 0-100m, 20 MAR 2001, A. Azofeifa, Mantillo L_S_269900_507750 #64457" (EMEC207846); 1 female, labeled: "COSTA RICA: Prov. Limon. R.B. Hitoy Cerere Send. a Espavel. 200m. 22 ENE 2003. W. Arana. Tp de Mantillo I. N. 184700 644200 #72874" (EMEC654899); 2 males and 1 female, labeled: "COSTA RICA: Puntarenas Reserva Biol. Carana 09°47'N, 84°36'W, 30m 23 July 1985, J. Longino #534-5, ex., leaf litter" (EMEC207995, EMEC207996, EMEC207997); 1 female, labeled: "COSTA RICA. Prov. Limon, Res Biol Hitoy Cerere, Est Hitoy Cerere, Send Bobocara. 300m. 11 FEB 2000. W. Arana Mantillo L_N_184250_641800 #56325" (EMEC207840); 1 female, labeled: "COSTA RICA. Prov. Limon, Central, Reserva Biol Hitoy Cerere, Est Hitoy Cerere. Send Tepezquintle 140m. 9-11 SET 1999. W. Arana Mantillo L_N_184250_643200 #56886" (EMEC207844); 1 female, labeled: "COSTA RICA. Prov. Limon R.B. Hitoy Cerere Sendero Tepezcuintle 0-100m.14 ENE 2003. W. Arana Tp. de Mantillo. I.N. 184200 643200 #72872" (EMEC654901); 1 female, labeled: "GUAT. Isabal Las Escobas 12.XI.1986 M. Sharkey" (EMEC654885); 1 male, labeled: "Ayutla, Guat. VIII-19-1947 B. Malkin" (EMEC654884); 1 male, labeled: "MEX. Chis., Palenque 28.VII.1983, rain forests, S. & J. Peck ultraviolet light" (EMEC654886). All specimens deposited in INBio except EMEC207991, EMEC207995, EMEC207997, EMEC207844 deposited in NMNH and EMEC207995, EMEC207846 deposited in EMEC.

Diagnosis.

This species most closely resembles Cratocerus indupalmensis and can be distinguished by the smaller size, differently shaped pronotum (Fig. 9e, f), and the differently sized and shaped median lobe of the male genitalia (Fig. 11d, e).

Description.

Size small for the genus, ABL 4.1-4.7 mm, TW 1.8-2.1 mm. Color. Head and prothorax brunneopiceous; elytra, abdominal sternites, and mandibles brunneous; legs, antennae, and remaining mouthparts rufotestaceous. Head. HL 0.5-0.7 mm, HW 0.6-0.8 mm. One supraorbital seta present over each eye. Apices of mandibles slightly curved, at less than 45 degree angle to mandibular midline. Apical maxillary palpomere length 6 × width at base. Mentum with triangular lateral lobes. Thorax. Pronotum (Fig. 9d) with distinct median impression, one distinct basal fovea laterad; one lateral seta located on apicolateral pronotal bead; one posterior seta present on posterior bead and projecting over posterior angle; pronotal anterior margin convex, greatest width just anterad of center; pronotal width 1.3-1.4 × pronotal length. Elytral length 1.3-1.5 × combined elytral width. Legs. Protibia with three distinct, stout spines confined to distal half of lateral margin; distance between spines 1 and 2 approximately half of distance between spines 2 and 3 (Fig. 10c). Genitalia, male. Median lobe curved sharply, between forty-five and ninety degrees, with respect to basal 1/3, remaining 2/3 with continuing curvature; not expanded apically; length short for genus. Left paramere simply curved (Figure 11d). Genitalia, female. Spermatheca with entire length thin, not gradually expanding towards apex, form similar to spermatheca of Cratocerus kavanaughi (Fig. 12c), but much longer (2 × length) and relatively narrower (1/2 × width).

Etymology.

This species is named in recognition of Tanya Shahjanian. As an undergraduate student at the University of California, Berkeley, Tanya participated in an insect art and culture project competition and was selected by the panel of judges as the winner with a sculpture piece called "It's an insect world". As a prize this fine species of beetle is named for her.

Distribution.

This species is distributed (Fig. 13) from Mexico through the Central American countries of Guatemala and Costa Rica.

Ecology.

Specimens have been collected from underneath decaying wood and at ultraviolet light traps from July to November.