Syntozyga Lower, 1901

Yang, Wenxu, Dong, Ruiqin, Song, Xueling & Yu, Haili, 2021, The genus Syntozyga Lower (Lepidoptera, Tortricidae) in China, with descriptions of two new species, ZooKeys 1028, pp. 95-111 : 95

Syntozyga Lower, 1901

Syntozyga Lower, 1901: 70. Type species: Syntozyga psammetalla Lower, 1901.

Eleuthodema Bradley, 1957: 95. Type species: Polychrosis pedias Meyrick, 1920.

Bubonoxena Diakonoff, [1968]: 40. Type species: Bubonoxena spirographa Diakonoff, [1968].

Cyclacanthina Diakonoff, 1973: 337. Type species: Cyclacanthina episema Diakonoff, 1973.

Remarks.

Superficially, adults of Syntozyga are similar to many species of Olethreutini , especially members of the subtribe Olethreutae, but they lack secondary sexual structures on the wing and legs in the male, that makes it easy to identify them from Olethreutae. In the male genitalia, Syntozyga is readily separated from most genera of Olethreutini by the absence of an uncus and by strongly reduced socii. In the Olethreutinae , reduction of the appendages of the tegumen is also found in the tribe Grapholitini , but their valvae are extremely simple in structure and evenly spined. In contrast, Syntozyga is characterized by a valva with a broad sacculus projecting distally and carrying varied vestiture such as spine clusters and strong bristles or thorns; sometimes a naked lobe (or two) is present between the cucullus and the apex of sacculus ( S. cerchnograpta Razowski, 2014, S. triangulana Aarvik, 2008, S. spirographa , S. similispirographa sp. nov., and S. apicispinata sp. nov.). Detailed descriptions of the morphology of Syntozyga were provided by Diakonoff (1968, 1973) and Horak (2006).