Eumacrocyrtus robertfoxi Cabras, 2024

Cabras, Analyn A., Buenavente, Perry Archival C. & Medina, Milton Norman, 2024, New species in the genera Eumacrocyrtus Schultze, 1923 and Enoplocyrtus Yoshitake, 2017 from Luzon Island, Philippines (Coleoptera, Curculionidae, Entiminae, Pachyrhynchini), ZooKeys 1191, pp. 23-33 : 23

Eumacrocyrtus robertfoxi Cabras sp. nov.

Figs 1-4 , 7-8 , 9

Type material.

Holotype (Figs 1 , 2 ), female: "Philippines- Luzon Island, Zambales, Villar / October, 1947/ leg. R.B. Fox (typed on white card) // HOLOTYPE female / Eumacrocyrtus robertfoxi sp. nov. / CABRAS, 2024 (typed on red card)" (PNM 15217).

Description.

Dimensions: LB: 13.5 mm. LR: 2.5 mm. WR: 1.2 mm. LP: 3.3 mm. WP: 3.8 mm. LE: 10.0 mm. WE: 6.0 mm. N = 1. Integument black. Body surface matte, rostrum, head, legs, and underside moderately lustrous. Head dorsal surface moderately rugose and irregularly punctured with few sparse appressed setae; lateroventral side below eye with sparse appressed setae; forehead between eyes flat with distinct midline groove reaching frons; eyes medium-sized, feebly convex, and weakly prominent on lateral outline of head. Rostrum dorsal surface moderately rugose on basal half with coarse punctures up to anterior two-thirds and finely punctate on apical third, and with sparse minute appressed setae; twice as long as wide (LR/WR:2.5 mm/1.2 mm); with distinct transverse and deep basal groove but not reaching lateral margin, dorsum with a distinct and narrowly shallow midline groove reaching basal half; dorsal contour flat until anterior two-thirds then gradually declining towards apex; lateral sides with subtruncate margin, wide from base then gradually constricted towards middle, and gradually widened towards apex. Antennal scape longer than funicle, scape reaching posterior margin of eye, sparsely covered with subappressed pubescence, and funicle with suberect brownish setae. Funicular segment I slightly longer than II, four times longer as wide, funicular segment II approximately three times longer as wide, segments III-V as long as wide, slightly shorter and narrower than VI and VII, segments VI as long as wide, slightly longer and wider than III-V and VII slightly wider than long, wider and longer than VI; club lanceolate.

Prothorax cylindrical, wider at base than apex, wider than long (LP/WP:3.3/3.8 mm), finely punctate and mostly glabrous with very few sparse setae, widest near base, weakly convex on dorsal surface, dorsal contour highest point at middle; with two dimple like depression on each side of disc. Elytra ovate, moderately longer than wide (LE/ WE:10.0/6.0 mm), three times as long and twice as wide as prothorax (WE/ WP: 6.0/3.8 mm, LE/LP: 10.0/3.3 mm), coarsely and irregularly punctate, with sparse minute pale-yellow to off-white appressed round scales on dorsum and dense yellow-ochre round scales on lateral sides, dorsum weakly convex, lateral sides near base and apex with round depressions, dorsal contour highest at middle, lateral contour gradually widening from basal margin towards middle then gently constricted towards apex and forming a mammilla-shaped apex, widest at middle, apex with brown erect setae. Legs with moderately clavate femora. Femora black covered with brown appressed setae. Fore tibiae covered with subappressed brown setae, inner edge moderately serrate with short denticles, and long brown suberect setae. Fore and mid tibiae bear a mucro at apex. Mid and hind tibiae covered with appressed brown setae, inner edge with brown dense suberect setae and few denticles along inner edge towards the apex. Tarsomeres pubescent. Fore coxae with sparse appressed golden yellow round, ovate and piliform scales towards anterior side, and sparse suberect metallic piliform scales towards posterior side. Mid and hind coxa with sparse appressed yellowish piliform scales. Mesoventrite with sparse appressed golden yellow piliform scales on disc and dense golden yellow piliform scales on distal ends. Metaventrite moderately depressed, especially at anterior margin, densely beset with golden yellow piliform scales on disc and thin ovate and piliform scales on distal ends. Ventrite I weakly depressed on disc, densely beset with golden yellow piliform scales on disc. Ventrite II weakly depressed and beset with golden yellow piliform scale on anterior half of disc. Ventrite III to V with fine sparse brown setae.

Male. Unknown.

Diagnosis.

The new species is different from the only known species Eumacrocyrtus canlaonensis Schultze, 1923 (Fig. 10 ) based on the following morphological differences: a) rostrum longer and more slender with a more angled dorsolateral edge near the base compared to E. canlaonensis with almost rounded edge (Figs 5-8 ), b) head with distinct and deep median furrow, and weak rugae, c) outline of head and rostrum discontinuous with a more distinct transverse basal groove reaching near the lateral margin (Figs 5-8 ), d) integuments matte black, e) elytra longer with coarser punctures, and rounded basal and post-median depression on lateral sides, and f) more slender and longer mammilla-shaped apex of elytra. In addition, the new species is found on Luzon Island outside the known range of E. canlaonensis which is only known from Negros Island (Fig. 21 ).

Etymology.

The species epithet " robertfoxi " is dedicated to Robert Bradford Fox (1918-1985), who collected the type material in Zambales for his groundbreaking and unparalleled discoveries in anthropology, which significantly advanced our current knowledge of the pre-Hispanic era from the Philippines.

Distribution.

Eumacrocyrtus robertfoxi sp. nov. is known only by the type specimen from Zambales, Luzon Island.

Figures

Figure 21. Distribution of E. robertfoxi sp. nov. (yellow), E. canlaonensis Schultze, 1923 (red) and Enoplocyrtus angelalcalai sp. nov. (blue) in the Philippines.

Figures 1 - 4. Eumacrocyrtus robertfoxi sp. nov., female holotype 1 dorsal habitus 2 lateral habitus 3 elytra, lateral view 4 meso-metasternum and abdominal sternites.

Figures 5 - 8. Head of Eumacrocyrtus spp. 5, 6 Eumacrocyrtus canlaonensis Schultze, 1923 5 dorsal view 6 lateral view 7, 8 Eumacrocyrtus robertfoxi sp. nov. 7 dorsal view 8 lateral view.

Figures 9, 10. Type specimens 9 Eumacrocyrtus robertfoxi sp. nov. 10 Eumacrocyrtus canlaonensis Schultze, 1923.