Silene nutans dubia (Herbich) Zapal ., Bull. Int. Acad. Sci. Cracovie, Cl. Sci. Math., Ser . B, Sci. Nat. 11: 151 (1911)

Novikov, Andriy, 2023, An annotated nomenclatural checklist of endemic vascular plants distributed in the Ukrainian Carpathians, Biodiversity Data Journal 11, pp. 103921-103921 : 103921

Silene nutans dubia (Herbich) Zapal., Bull. Int. Acad. Sci. Cracovie, Cl. Sci. Math., Ser. B, Sci. Nat. 11: 151 (1911)

Silene dubia ≡ Silene dubia Herbich, Fl. Bucovina: 388 (1859), non alior; GBIF: https://www.gbif.org/species/7592009; POWO: https://powo.science.kew.org/taxon/2488403-4; JACQ: https://www.jacq.org/detail.php?ID=442588; JACQ: https://www.jacq.org/detail.php?ID=442591; JACQ: https://www.jacq.org/detail.php?ID=442592; JACQ: https://www.jacq.org/detail.php?ID=442595; JACQ: https://www.jacq.org/detail.php?ID=442597; JSTOR Global Plants: https://plants.jstor.org/stable/10.5555/al.ap.specimen.cher0200022; JSTOR Global Plants: https://plants.jstor.org/stable/10.5555/al.ap.specimen.cher0200023; JSTOR Global Plants: https://plants.jstor.org/stable/10.5555/al.ap.specimen.cher0200019; JSTOR Global Plants: https://plants.jstor.org/stable/10.5555/al.ap.specimen.cher0200020; JSTOR Global Plants: https://plants.jstor.org/stable/10.5555/al.ap.specimen.cher0200021

Silene dubia ≡ Silene dubia Herbich ex Rohrb., Monogr. Silene : 217 (1869), non alior; GBIF: https://www.gbif.org/species/8297181; IPNI: https://www.ipni.org/n/157265-1; WFO: http://www.worldfloraonline.org/taxon/wfo-0000440631; BHL: https://www.biodiversitylibrary.org/page/15321329#page/229

Silene nutans ≡ Silene nutans var. dubia (Herbich) Zapał., Sprawozd. Kom. Fizjograf. 24: 111 (1889)

Silene dubia = Silene dubia var. glabriuscula ( Zapał.) Guşul., Fl. Rep. Pop. Rom. 2: 179 (1953)

Silene dubia = Silene dubia var. hormuzakii Guşul., Fl. Rep. Pop. Rom. 2: 179 (1953)

Silene dubia = Silene dubia var. hormuzakii f. acaulis Guşul., Fl. Rep. Pop. Rom. 2: 179, 665 (1953) [monster forma]

Silene dubia = Silene dubia var. hormuzakii f. apricorum ( Zapał.) Graebn. in Asch. & Graebn. emend. Guşul., Fl. Rep. Pop. Rom. 2: 179 (1953)

Silene dubia = Silene dubia var. hormuzakii f. herbichii ( Zapał.) Graebn. in Asch. & Graebn. emend. Guşul., Fl. Rep. Pop. Rom. 2: 179 (1953)

Silene dubia = Silene dubia var. hormuzakii f. kelemenensis ( Zapał.) Graebn. in Asch. & Graebn. emend. Guşul., Fl. Rep. Pop. Rom. 2: 179 (1953)

Silene dubia = Silene dubia var. hormuzakii f. lilacina ( Zapał.) Guşul., Fl. Rep. Pop. Rom. 2: 179 (1953)

Silene dubia = Silene dubia var. hormuzakii f. robustior (Schur) Graebn. in Asch. & Graebn. emend. Guşul., Fl. Rep. Pop. Rom. 2: 179 (1953)

Silene nutans = Silene nutans subsp. dubia var. dubia f. apricorum Zapał., Consp. Fl. Galic. Crit. 3: 195 (1911)

Silene nutans = Silene nutans subsp. dubia [unranked] b Melampyrum herbichii Zapał., Consp. Fl. Galic. Crit. 3: 195 (1911)

Silene nutans = Silene nutans subsp. dubia [unranked] a kelemenensis Zapał., Consp. Fl. Galic. Crit. 3: 195 (1911)

Silene nutans = Silene nutans subsp. dubia [unranked] a kelemenensis f. lilacina Zapał., Consp. Fl. Galic. Crit. 3: 195 (1911)

Silene nutans = Silene nutans subsp. dubia var. dubia f. luxuriosa Zapał., Consp. Fl. Galic. Crit. 3: 195 (1911)

Silene nutans = Silene nutans subsp. dubia var. dubia f. tenuis Zapał., Consp. Fl. Galic. Crit. 3: 195 (1911)

Silene nutans = Silene nutans [unranked] c. Silene glabriuscula Zapał., Consp. Fl. Galic. Crit. 3: 192 (1911); GBIF: https://www.gbif.org/species/11954760

Silene nutans = Silene nutans [unanked] β Silene transsilvanica Grec., Consp. Fl. Rom.: 109 (1898)

Silene transsilvanica = Silene transsilvanica Schur, Oesterr. Bot. Z. 8: 22 (1858) [nom. nudum] et Oesterr. Bot. Z. 10: 181 (1860); GBIF: https://www.gbif.org/species/5587908; IPNI: https://www.ipni.org/n/158548-1; WFO: http://www.worldfloraonline.org/taxon/wfo-0000440073; POWO: https://powo.science.kew.org/taxon/158548-1; BHL: https://www.biodiversitylibrary.org/page/28724891#page/30; JACQ: https://www.jacq.org/detail.php?ID=369397; JACQ: https://www.jacq.org/detail.php?ID=369398; JACQ: https://www.jacq.org/detail.php?ID=369401; JACQ: https://www.jacq.org/detail.php?ID=369407; JACQ: https://www.jacq.org/detail.php?ID=369408; JACQ: https://www.jacq.org/detail.php?ID=369417; JSTOR Global Plants: https://plants.jstor.org/stable/10.5555/al.ap.specimen.lw00204020; JSTOR Global Plants: https://plants.jstor.org/stable/10.5555/al.ap.specimen.lw00204018a; JSTOR Global Plants: https://plants.jstor.org/stable/10.5555/al.ap.specimen.lw00204022; JSTOR Global Plants: https://plants.jstor.org/stable/10.5555/al.ap.specimen.lw00204019; JSTOR Global Plants: https://plants.jstor.org/stable/10.5555/al.ap.specimen.lw00204018b; JSTOR Global Plants: https://plants.jstor.org/stable/10.5555/al.ap.specimen.lw00204017

Silene transsilvanica = Silene transsilvanica var. angustifolia Hormuz., Oesterr. Bot. Z. 61: 147 (1911); BHL: https://www.biodiversitylibrary.org/item/37201#page/159

Silene saxatilis = Silene saxatilis [unranked] a Silene saxatilis racemosa Schur, Enum. Pl. Transsilv.: 101 (1866); BHL: https://www.biodiversitylibrary.org/item/7364#page/121

Silene saxatilis = Silene saxatilis [unranked] a Silene saxatilis robustior Schur, Enum. Pl. Transsilv.: 101 (1866); BHL: https://www.biodiversitylibrary.org/item/7364#page/121; JSTOR Global Plants: https://plants.jstor.org/stable/10.5555/al.ap.specimen.lw00203992

Silene saxatilis - Silene saxatilis Schur, Enum. Pl. Transsilv.: 101 (1866), non Sims nec M.Bieb.; BHL: https://www.biodiversitylibrary.org/item/7364#page/121

Conservation status

In Ukraine - LC ( Onyshchenko et al. 2022).

Distribution

SE Carpathian endemic.

Notes

Following Euro+Med (https://europlusmed.org/cdm_dataportal/taxon/d201548b-883c-4f46-8151-aed4fac379ee, accessed on 05.06.2023) and Worldplants (https://www.worldplants.de/world-plants-complete-list/complete-plant-list?name=Silene-nutans, accessed on 07.06.2023), S. nutans L. includes three subspecies - subsp. nutans (most widely distributed, with Eurasian range), subsp. insubrica (Gaudin) Soldano (distributed mainly in the Alps, but also occurs in Greece, Hungary, Romania and some other countries) and subsp. Scabiosa nutans dubia (occurs exclusively in the south-eastern Carpathians). Additionally, S. nutans subsp. smithiana (Moss) Jeanm. & Bocquet distributed in France, Belgium, Great Britain and the Netherlands was described by Jeanmonod and Bocquet (1983). Silene nutans subsp. smithiana in mentioned databases is synonymised with S. nutans susbp. Silene nutans nutans . However, recent studies showed that this subspecies is a separate taxon represented by two haplotypes ( Martin et al. 2016). Besides this, POWO (https://powo.science.kew.org/taxon/157927-1, accessed on 05.06.2023) and WFO (https://list.worldfloraonline.org/wfo-0000440799, accessed on 05.06.2023) recognise S. nutans subsp. livida (Willd.) Gremli. However, S. nutans subsp. livida is a direct synonym of S. nutans subsp. insubrica ( Jeanmonod and Bocquet 1983, Soldano 1991, Soldano 2001).

Two of the four mentioned subspecies occur in the Ukrainian Carpathians ( Chopyk and Fedoronchuk 2015) - S. nutans susbp. Silene nutans nutans (occupies light forested and adjacent habitats in the montane belt) and S. nutans subsp. dubia (occupies rocky and stony slopes in the montane and subalpine belts).