Heteropachylus, ROEWER, 1913

Mendes, Amanda Cruz, 2011, Phylogeny and taxonomic revision of Heteropachylinae (Opiliones: Laniatores: Gonyleptidae), Zoological Journal of the Linnean Society 163 (2), pp. 437-483 : 466

HETEROPACHYLUS ROEWER, 1913

Heteropachylus Roewer, 1913a: 75, 1923: 421 , 1929: 185; Mello-Leitão, 1923: 123, 1926: 342, 1932: 185, 1935: 100; Kästner, 1937: 389; Soares & Soares, 1954: 267, 1974: 611. H. Soares, 1968: 140; Muñoz-Cuevas, 1973: 226; Kury, 2003a: 142.

Aesotrinoma Soares, 1977: 251 ; Kury, 1994: 350; 2003a: 142 (type species = Aesotrinoma crassicalcanei Soares, 1977 , by original designation). syn. nov.

Chavesincola Soares & Soares, 1946: 238, 1954: 243 ; Kury, 1994: 350, 2003a: 142. syn. nov.

Distribution ( Fig. 28 ): BRAZIL. Espírito Santo and Bahia. WWF Ecoregions: NT0103 and NT0104, respectively, Bahia coastal forests and Bahia interior forests.

Type species: Heteropachylus spiniger Roewer, 1913 , by monotypy.

Diagnosis: Three or four mesotergal areas; mesotergal area III with variable armature. Median armature of free tergite II with length comparable to the length of the tergite ( Figs 15A , 17A , 19A , 21A , 23A ). Calcaneus I about half the size of the metatarsus and lighter than the astragalus. Femur IV ( Figs 15 , 18 , 20 , 22 , 24 ) distinctly curved in dorsal view, with two curvatures (‘S-shaped’) in lateral view; bearing dorsobasal apophysis and row of tubercles on the prolateral margin, rounded at the distal half, larger at the medial portion of the row. Male genitalia ( Fig. 25 ). Dorsal digitiform process in glans mostly absent (except in H. crassicalcanei ).

Figures

Figure 15. Heteropachylus inexpectabilis comb. nov. A, C–E, male MNRJ. A, habitus, dorsal view; C, frontal view; D,habitus, lateral view; E, ventroposterior view. B, female MNRJ, habitus, dorsal view. Scale bars = 1 mm.

Figure 17. Heteropachylus crassicalcanei comb. nov. Male holotype MNRJ 5216. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, patella and apex of femur IV. Scale bars = 1 mm.

Figure 18. Heteropachylus crassicalcanei comb. nov. Male holotype MNRJ 5216. Right trochanter, femur and patella IV. A, dorsal view. B, ventral view. C, prolateral view. D, retrolateral view. Scale bars = 1 mm.

Figure 19. Heteropachylus gracilis sp. nov. A, C–F, male holotype IBSP440. A, habitus, dorsal view; C, frontal view; D, patella and apex of femur IV; E, habitus, lateral view; F, ventroposterior view. B, female paratype IBSP 450. Habitus, dorsal view. Scale bars = 1 mm.

Figure 20. Heteropachylus gracilis sp. nov. Male holotype IBSP440. A, left trochanter, femur, and patella IV, dorsal view. B, C, right trochanter, femur, and patella IV. B, ventral view; C, prolateral view. D, left trochanter, femur, and patella IV, retrolateral view. Scale bars = 1 mm.

Figure 21. Heteropachylus peracchii comb. nov. A, C–F, male holotype HEMS 485. A, habitus, dorsal view; C, frontal view; D, patella and apex of femur IV; E, habitus, lateral view; F, ventroposterior view. B, female HEMS 665. Habitus, dorsal view. Scale bars = 1 mm.

Figure 22. Heteropachylus peracchii comb. nov. Male holotype HEMS 485. Left trochanter and femur IV. A, dorsal view. B, ventral view. C, prolateral view. D, retrolateral view. Scale bars = 1 mm.

Figure 23. Heteropachylus spiniger. Male lectotype SMF RI 798. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, apical view of right femur IV. Scale bars = 1 mm.

Figure 24. Heteropachylus spiniger. Male lectotype SMF RI 798. A, right trochanter, femur, and patella IV. Dorsal view. B, right trochanter and femur. Ventral view. C, right femur and patella IV. Prolateral view. The arrow indicates the anvil-shaped dorsobasal apophysis. D, right femur and patella IV. Retrolateral view. The arrow indicates the two subdistal tubercles. Scale bars = 1 mm.

Figure 25. Male genitalia of species of Heteropachylus. A, B, Heteropachylus ramphonotus sp. nov. holotype IBSP 994. A, dorsal view; B, lateral view. C, D, Heteropachylus inexpectabilis comb. nov. MNRJ. C, dorsal view; D, lateral view. E, F, Heteropachylus crassicalcanei comb. nov. MNRJ 5216. E, dorsal view; F, lateral view. G, H. Heteropachylus gracilis sp. nov. paratype IBSP 450. G, dorsal view; H, lateral view. I, J, Heteropachylus peracchii HEMS 665. I, dorsal view; J, lateral view. K, L, Heteropachylus spiniger lectotype SMF RI 798. K, dorsal view; L, lateral view. Scale bars = 0.1 mm.

Figure 28. Geographical distribution of the species of Heteropachylus.