Fragosa rugosifrons (Schiner, 1868) Miranda & Skevington & Marshall, 2020

Miranda, Gil Felipe Gonçalves, Skevington, Jeffrey H. & Marshall, Stephen A., 2020, New generic concepts for orphaned lineages formerly treated as part of the genus Ocyptamus Macquart, 1834 (Diptera, Syrphidae), Zootaxa 4822 (2), pp. 151-174 : 154

Fragosa rugosifrons species group (Fragosa sensu stricto)

Description. Head. Face usually dark dorsal to tubercle; tubercle pointed and medially positioned ( Fig. 5 ). Frons dark. Frons/frontal triangle rugose ( Fig. 4 ). Frontal triangle with white microtrichosity concentrated laterally along eye margin ( Fig. 83 ). Female ocellar triangle separated by less than its length from posterior eye margin ( Fig. 1 ). Occiput microtrichia homogeneously distributed ( Fig. 5 ). Thorax. Scutum dull pale-microtrichose with sub-median pair of weak white microtrichose patches on anterior region, sometimes with weak stripes in place of patches ( Fig. 1 ). Scutellum pale. Post-metacoxal bridge almost complete ( Fig. 79 ). Metafemur pale. Wing. Female wing hyaline; female wing bare on basal 1/4 (or almost the whole cell) of cell c, basal 1/3 to 1/2 of br, and baso-anterior margin of bm. Terminalia. Male epandrium enlarged; hypandrium reduced; *surstylus quadrangular with filiform apical extension ( Fig. 7 ); *subepandrial sclerite with strongly sclerotized rectangular middle area and wide, but weakly sclerotized, ventrally directed lateral expansions ( Fig. 9 ); *hypandrium oval ( Fig. 11 ), much smaller than epandrium; postgonite elongated ( Figs 10, 11 ).

Included species (11): F. argentina ( Curran, 1939) comb. nov. [1b], F. deceptor ( Curran, 1930a) comb. nov. [1b], F. filiola ( Shannon, 1927) comb. nov. [2], F. harlequina ( Hull, 1948) comb. nov. [1b, 1c], F. hyacinthia ( Hull, 1947a) comb. nov. [2], F. mara ( Curran, 1941) comb. nov. [1b], F. oenone ( Hull, 1949a) comb. nov. [2, type lost], F. provocans ( Curran, 1939) comb. nov. [1b], F. rugosifrons ( Schiner, 1868) , comb. nov. [2, 3, 4], F. stenogaster ( Williston, 1888) comb. nov. [1b], F. zephyrea ( Hull, 1947b) comb. nov. [1b].

Comments. The rugose frons/frontal triangle is a distinct character of this group ( Fig. 4 ), but not unique since it occurs in the Pelecinobaccha brevipennis species group as well. Further distinct characters for the F. rugosifrons group include the face that is dark dorsal to tubercle, the pointed and medially positioned facial tubercle ( Fig. 5 ), the entirely pale scutellum, the almost complete post-metacoxal bridge ( Fig. 79 ), the enlarged epandrium ( Fig. 7 ), and the reduced hypandrium ( Fig. 10 ).

Figures

FIGURES 1–11. Fragosa Miranda gen. nov. 1: Fragosa sp. (INPA-DIP001362), female, habitus, dorsal view, photo previously published in Mengual et al. (2018). 2: Fragosa aff. titania (CNC), female, abdomen, dorsal view, photo previously published in Mengual et al. (2018). 3: Fragosa argentina (Curran) (CNC_Diptera196006), female, wing. 4: Fragosa stenogaster (Williston) (debu01088839), female, head, frontal view, photo previously published in Mengual et al. (2018). 5: Fragosa stenogaster (Williston) (debu01088840), male, head, lateral view. 6: Fragosa sp., female, habitus, dorsal view, by Steve Marshall. 7: Fragosa stenogaster (Williston) (debu01088840), male, epandrium, lateral view, photo previously published in Mengual et al. (2018). 8: Fragosa stenogaster (Williston) (debu01088839), female, apex of abdomen, dorsal view, photo previously published in Mengual et al. (2018). 9–11. Fragosa stenogaster (Williston) (debu01088840), male. 9: epandrium, ventral view. 10: hypandrium, lateral view. 11: hypandrium, ventral view. ap: apodeme; b: basiphallus; bm: cell bm; br: cell br; c: cell c; d: distiphallus; e: epiproct; f: frons; h: hypandrium; oc: occiput; p: postgonite; s: surstylus; se: subepandrial sclerite; t: facial tubercle; t2, t3, t7 and t8: terga 2, 3, 7 and 8.

FIGURES 78–88. 78: Fragosa harlequina (Hull) (holotype Baccha harlequina Hull, AMNH), male, thorax, lateral, by AMNH (line drawing represents outline of the alula). 79: Fragosa argentina (Curran) (CNC_Diptera174232), thorax, postero-ventral (line drawing represents outline of the incomplete post-metacoxal bridge). 80–81: Fragosa virgilio (Hull) (holotype Baccha virgilio Hull, CNC, CNC_Diptera189124), male. 80: habitus, lateral, by CNC. 81: head, oblique frontal, by CNC. 82: Fragosa titania (Hull) (JSS25229), female, head, dorsal. 83: Fragosa stenogaster (Williston) (debu01088840), male, head, dorsal. 84–85: Hypocritanus fascipennis (Wiedemann), female, head. 84: frontal, by Ximo Mengual (http://syrphidae.myspecies.info/ taxonomy/term/879). 85: lateral (CNC_Diptera501). 86–87: Nuntianus cecrops (Hull) (INPA-DIP000246), female, head. 86: dorsal. 87: lateral. 88: Victoriana aff. melanorrhina (DEBU), female, head, lateral. a: alula; ai: antennal insertion; f: frons; ft: frontal triangle; lem: lateral eye margin; mt: metafemur; o: ocellar triangle; oc: occiput; pmt: post-metacoxal bridge; t: facial tubercle.