Fragosa tenuis (Walker, 1852) Miranda & Skevington & Marshall, 2020

Miranda, Gil Felipe Gonçalves, Skevington, Jeffrey H. & Marshall, Stephen A., 2020, New generic concepts for orphaned lineages formerly treated as part of the genus Ocyptamus Macquart, 1834 (Diptera, Syrphidae), Zootaxa 4822 (2), pp. 151-174 : 154

Fragosa tenuis species group

Description. Head. Face wholly pale; tubercle convex and ventrally positioned ( Fig. 81 ). Frons narrow (less than 1/3 of head width) and pale on ventral 1/4 to 1/2 lateral to lunule ( Fig. 82 ). Frons/frontal triangle smooth. Frontal triangle white microtrichia homogeneously distributed but oriented ventro-dorsally (only visible from a ventral view) ( Fig. 81 ). Female ocellar triangle separated by more than its length from the posterior eye margin ( Fig. 82 ). Dorsal occiput with microtrichia oriented ventro-dorsally ( Figs 80, 82 ). Dorsal occiput with 1–2 rows of pile ( Fig. 82 ). Thorax. Scutum shiny black. Metathoracic epimera extended dorsal to the metacoxa but not so close to each other as in the F. rugosifrons group ( Fig. 79 ). Metafemur with sub-apical dark band ( Fig. 80 ). Wing. Female wing hyaline basally and gradually turning light brown towards apex; female wing bare on basal 1/3 to 2/3 of c, and bases of br and bm. Terminalia. Male epandrium and hypandrium of similar size ( Figs 92, 94 ); surstylus sub-quadrangular with slightly extended apex ( Fig. 93, 94 ); subepandrial sclerite rectangular and wide, with basal corners slightly extended and round ( Fig. 95 ).

Included species (6): F. aurora ( Hull, 1943c) comb. nov. [1b], F. filissima ( Hull, 1943a) comb. nov. [1a], F. macer ( Curran, 1930a) comb. nov. [1b], F. tenuis ( Walker, 1852) comb. nov. [2, type lost], F. titania ( Hull, 1943a) comb. nov. [1b], F. virgilio ( Hull, 1942b) comb. nov. [1a].

Comments. Fragosa aurora ( Fig. 74 ) and F. virgilio ( Fig. 76 ) were previously allocated to the O. lepidus group ( Mengual et al. 2018). However, the type specimen of F. virgilio and photos of the type and the original description of F. aurora were studied and they show the distinct very narrow and long 2 nd abdominal segment that defines the genus Fragosa , and the pale smooth frontal triangle that defines the F. tenuis group.

Figures

FIGURES 78–88. 78: Fragosa harlequina (Hull) (holotype Baccha harlequina Hull, AMNH), male, thorax, lateral, by AMNH (line drawing represents outline of the alula). 79: Fragosa argentina (Curran) (CNC_Diptera174232), thorax, postero-ventral (line drawing represents outline of the incomplete post-metacoxal bridge). 80–81: Fragosa virgilio (Hull) (holotype Baccha virgilio Hull, CNC, CNC_Diptera189124), male. 80: habitus, lateral, by CNC. 81: head, oblique frontal, by CNC. 82: Fragosa titania (Hull) (JSS25229), female, head, dorsal. 83: Fragosa stenogaster (Williston) (debu01088840), male, head, dorsal. 84–85: Hypocritanus fascipennis (Wiedemann), female, head. 84: frontal, by Ximo Mengual (http://syrphidae.myspecies.info/ taxonomy/term/879). 85: lateral (CNC_Diptera501). 86–87: Nuntianus cecrops (Hull) (INPA-DIP000246), female, head. 86: dorsal. 87: lateral. 88: Victoriana aff. melanorrhina (DEBU), female, head, lateral. a: alula; ai: antennal insertion; f: frons; ft: frontal triangle; lem: lateral eye margin; mt: metafemur; o: ocellar triangle; oc: occiput; pmt: post-metacoxal bridge; t: facial tubercle.

FIGURES 89–102. 89: Victoriana sativa (Curran) (holotype Baccha sativa Curran, AMNH), habitus, dorsal, by AMNH. 90– 91: Victoriana laudabilis (Williston) (holotype Baccha laudabilis Williston, BMNH, NHMUK010369870), copyright of The Trustees of the Natural History Museum, London. 90: base of abdomen and metafemur, dorsal. 91: base of wing, dorsal (line drawing represents outline of the alula). 92–95: Fragosa sp. (species group tenuis, CNC_Diptera196012), male. 92: hypandrium, lateral. 93: surstylus, ventro-lateral. 94: epandrium, lateral. 95: epandrium, ventral (line drawing represents outline of the subepandrial sclerite). 96–97: Nuntianus abata (Curran) (CNC_Diptera209361), male, hypandrium. 96: lateral. 97: ventral. 98: Syrphus conjunctus Wiedemann (currently incertae sedis, CNC_Diptera209347), male, hypandrium, lateral. 99: Victoriana aff. melanorrhina (debu00279441), male, hypandrium, ventral. 100: Victoriana aff. melanorrhina (DEBU), female, terminalia, dorsal. 101: Victoriana parvicornis (Loew) (CNC_Diptera237922), terminalia, dorsal. 102: Maiana pumila (Austen) (debu00178088), female, terminalia, dorsal. a: alula; c: cercus; d: distiphallus; e: epiproct; l: lobe; p: baso-posterior pile; t7 and 8: tergum 7 and 8.

FIGURES 68–77. 68: Maiana callida (Hine) (CNC_Diptera105396), male, abdomen, ventral. 69: Maiana pumila (Austen) (JSS32861), male, terminalia, oblique fronto-ventral. 70–73: Hybobathus pola (Curran). 70: female terminalia (CNC_Diptera 161361), apex, dorsal. 71–72: male terminalia (CNC_Diptera 161366). 71: hypandrium, ventral. 72: terminalia, lateral. 73: holotype Baccha pola Curran (AMNH), male, habitus, dorsal, by AMNH. 74: Fragosa aurora (Hull) (holotype Baccha aurora Hull, AMNH), male, habitus, dorsal, by AMNH. 75: Nuntianus variegatus (Macquart) (holotype Baccha variegata Macquart, MNHNP, ED8265), female, habitus, dorsal, by MNHNP. 76: Fragosa virgilio (Hull) (holotype Baccha virgilio Hull, CNC, CNC_Diptera189124), male, habitus, dorsal, by CNC. 77: Victoriana oblonga (Walker) (holotype Baccha oblonga Walker, BMNH, NHMUK 010369937), female, habitus, dorsal, copyright of The Trustees of the Natural History Museum, London. c: cercus; e: epiproct; h: hypandrium; l: lobe; n: lateral notch; p: postgonite; ph: phallus; s: surstylus; s4 and 5: sternum 4 and 5; t7: tergum 7.