Tomarus villosus (Burmeister, 1847)

Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36 : 13

Distribution. Tomarus villosus occurs in southern Peru, Chile, and Argentina.

Locality Records ( Fig. 17d ). 208 specimens examined from BCRC, CJME, CMNC, FSCA, LEULS, MACN, MNHN, MNNC, MLPA, UMSP, USNM.

ARGENTINA (17). NEQUÉN (17): Collón Cura, Quilquihue, Rinconada, Zapala.

CHILE (191). ANCONCAGUA (1): Los Molles. ANTOFAGASTA (1): San Pedro de Atacama. ARAU- CANÍA (8): Cautín-Temuco. ARICA Y PARINACOTA (6): Luta-Molino, Arica. ATACAMA (6): Caldera, Copiapó, Río Pulida, San Pedro de Atacama, Totoral-Copiaco. BIOBIO (7): Quillón. CHOAPA (1): Los Vilos. CONCEPCIÓN (3): Concepción. COQUIMBO (42): Choapa, Diaguita-Valle Elqui, Guampulla, Guanaquero, Huanta-Elqui, Llano Compañía-La Serena, La Serena, Limarí, Los Choros, Monte Grande- Elqui, Pueblo Diaguitas-Elqui, Punta Teatinos-La Serena, Salamanca, Tongoy, Totoralillo, Vicuña-Elqui. HUASCO (1): Vallenar (15 km N). LOS RÍOS (1): Reserva Costera Valdiviana. MAULE (31): Chance, Cuaquemes, Curanipe, Curicó (12 km SW), El Coigo, Los Quenes (6 km S), Pelluhue, Potrero Grande (35 km SE), Río Teno, San Javier, Suazal. ÑUBLE (3): Recinto, Rio Pinto E. of Chilican. O’HIGGINS (10): Pichilemu, Rosario. REGIÓN METROPOLITANA (32): Alhue, Cordillera Reserva Río Clarillo, Cuerta lo Prado-Santiago, El Toyo, San Cristóbal, Santiago. SANTIAGO (3): Pilay, Renca. TARAPACÁ (3): Iquique, Refresco-Tomarugal. VALPARAISO (14): Canelo, El Convento, El Quisco-San Antonio, Mantagua. NO DATA (18).

Temporal Distribution. January (24), February (24), March (8), May (5), July (5), August (9), September (10), October (14), November (18), December (15).

Diagnosis. Tomarus villosus is easily recognized because the teeth of the clypeal apex are subcontiguous ( Fig. 4h ), and the pronotal surface is nearly smooth or with micropunctures and lacking a subapical tubercle ( Fig. 8h ).

Life History. Adults are attracted to lights at night. Some specimens have been taken in coastal sand dunes near the ocean.

Figures

Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.

Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.

Figure 17. Distribution of Tomarus spp. a) T. roigjunenti n. sp. b) T. rosettae. c) T. spinipenis n. sp. d) T. villosus.