Tomarus maimon, Erichson, 1847

Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36 : 8

Distribution. Tomarus maimon is widely distributed in the low, arid coastal areas of Peru and Chile.

Locality Records ( Fig. 16c ). 55 specimens examined from CJME, CMNC, IADIZA, MNNC. CHILE (55). ARICA (39): Arica- Iquique, Valle de Zapala. (16): Tarapacá.

Temporal Distribution. January (2), February (3), October (5), November (10), December (36).

Diagnosis. Tomarus maimon is externally similar to T. gyas . The labial apex in T. gyas is narrowly notched at its center ( Fig. 7c ), while in T. maimon the apex of the labium is slightly convex ( Fig. 7d ). The tubercles on the head of T. gyas are rounded and widely separated from each other ( Fig. 4c ), whereas the tubercles of T. maimon are distinctly transverse and slightly less separated ( Fig. 4d ). In addition, the pronotal fovea ( Fig. 8c ) of T. gyas is larger than that of T. maimon ( Fig. 8d ). The parameres are different between both species (compare Fig. 11d , 12d , 13d and 11c , 12c , 13c ). Tomarus gyas has five copulatory lamellae on the internal sac ( Fig. 15c ), but in T. maimon there are three copulatory lamellae ( Fig. 15d ).

Life History. Adults are attracted to lights at night. Some specimens have been taken in sand dunes adjacent to ocean beaches.

Figures

Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.

Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.

Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.

Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.

Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.

Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.

Figure 15. Internal sac of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (AL) accessory lamella, (CL) copulatory lamella (CL), (Ra) raspula, (Esp) speculum, and (Fg) a complex, spike-like flagellum that varies in length and thickness.

Figure 16. Distribution of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. maimon. d) T. gyas.