Chydaeus loeffleri ( Jedlička, 1966 )

Kataev, Boris M. & Schmidt, Joachim, 2019, New data on the irvinei group of Chydaeus of the Himalaya from Nepal and southern Tibet (Coleoptera, Carabidae, Harpalini), Zootaxa 4686 (2), pp. 202-214 : 206-207

Chydaeus loeffleri ( Jedlička, 1966) , stat. rest.

( Figs 3, 4 , 13–30 )

Harpalus loeffleri Jedlička, 1966: 238

Diagnosis. In its combination of characters, including left mandible truncate at apex and elytra not fused along suture, Ch. loeffleri is most similar to Ch. irvinei , but slightly larger (length 6.8–7.8 mm, width 3.0– 3.4 mm), dorsal microsculpture developed throughout, distinct, pronotal basal angle with a tiny acute denticle protruding laterally, elytral sutural angle narrowly rounded just at tip, metacoxa with or without an additional, somewhat large medial setigerous pore, male pro- and especially mesotarsi less widened, protarsus with only tarsomeres 2–4 and mesotarsus with only tarsomeres 2 and 3 bearing adhesive vestiture, median lobe of aedeagus relatively smaller (MLL/L 0.21–0.24 versus 0.31–0.32), slightly concave along ventral margin (lateral view) and with a wider spiny patch at middle of median lobe.

Distribution ( Fig. 38 , ‘red circle’ and ‘red square’). Upper sources of the Tama Koshi River in the Greater Himalaya of eastern Central Nepal and southern Tibet: Rolwaling Valley (= Himalayan longitudinal Valley in eastern Central Nepal), and Rongshar valley (= uppermost portion of the Tama Koshi Valley in southern Tibet).

Remarks. The species forms two subspecies in two geographically adjacent valley systems.

Figures

FIGURE 38. Chydaeus, distribution: ‘red star’, Ch. irvinei (Andrewes. 1930); ‘red circle’, Ch. loeffleri loeffleri (Jedlička, 1966); ‘red square’, Ch. loeffleri tibetanus ssp. n.; ‘yellow square’, Ch. cf. loeffleri, population from Bothe Koshi Valley, see text for details; ‘blue circle’, Ch. soluensis sp. n.

FIGURES 1–5. Chydaeus, habitus (males). 1, Ch. irvinei (Andrewes. 1930), lectotype; 2, same, paralectotype from Gautsa; 3, Ch. loeffleri tibetanus ssp. n., holotype; 4, Ch. loeffleri loeffleri (Jedlička, 1966), specimen from the locality “Beding to Na”; 5, Ch. soluensis sp. n., paratype from Lamjura La. Scale bars = 1 mm.

FIGURES 13–25. Chydaeus loeffleri loeffleri (Jedlička, 1966). 13, apex of left mandible; 14–17, 19, 20, median lobe of aedea- gus (14, 16, 19, lateral view; 15, 17, 20, dorsal view); 18, terminal lamella, dorsal view; 21, apical stylomere, lateral view; 22, laterotergite and stylus, ventral view; 23, male protarsus, 24, male mesotarsus; 25, left metacoxa. 14, 15, holotype; 13, 16–25, specimens from the Rolwaling Valley. Abbreviations: mp, medial pore; pmp, posteromedial pore. Scale bars: A = 0.5 mm (Figs 13–24), B = 1 mm (Fig. 25).

FIGURES 26–30. Chydaeus loeffleri tibetanus ssp. n., holotype. 26, terminal lamella of aedeagus, dorsal view; 27, median lobe of aedeagus, lateral view; 28, same, dorsal view; 29, male protarsus; 30, male mesotarsus. Scale bar = 0.5 mm.