Chimarra tipulida, Johanson, Kjell Arne & Oláh, János, 2012

Johanson, Kjell Arne & Oláh, János, 2012, Revision of the Fijian Chimarra (Trichoptera, Philopotamidae) with description of 24 new species, Zootaxa 3354, pp. 1-58 : 49-50

Chimarra tipulida , new species

Figs. 28 , 150–154

Chimarra tipulida resembles C. psychodida , C. malickyi , and C. levuensis in having gonopods each with both a slender, strongly produced dorsal branch, and a shorter, well-defined, plate-like ventral lobe. It is distinguished from C. malickyi by having segment IX with a ventral margin that is more uniformly convex in lateral view, and in having a shorter ventral branch of each gonopod. It is separated from C. levuensis by having parallel lateral branches of tergum X and a longer ventral branch of each gonopod. It is separated from C. psychodida by having a more strongly produced anterior lateral lobe of segment IX, and the absence of a dorsal process at mid-length on the lateral branches of tergum X.

Male. Body pale brown, dorsal part of meso and metathorax brown. Large dark area between lateral and anterior ocelli. Foreleg anterior claw as long as foreleg spur.

Wings ( Fig. 28 ). Forewings 4.9 mm (n=1), greyish brown; broad, ratio of length to breadth 3.1; R1 slightly curved before crossvein r; radial sector weakly produced anterad immediately before discoidal cell; discoidal cell originating immediately before mid-length of wing, nearly 3x longer than wide; median cell slightly shorter than discoidal cell; crossvein r situated at base of fork I; fork I originating before crossvein s at distance equal to length of crossvein s; nygma located near base of fork II; fork III 1 /5th as long as wing; fork V as long as fork I, shorter than fork II; Cu2 well-separated from A at wing margin. Hind wings 3.9 mm (n=1), brown; broad, ratio of length to breadth 3.0; margin slightly incurved at arculus, where Cu1 and Cu2 fused with margin; fork III clearly longer than discoidal cell and 1/7th as long as wing; fork V slightly shorter than fork I; 1A+2A 4x longer than 1A.

Male genitalia ( Figs. 150–154 ). Segment IX clearly shorter than high; anterior plate nearly triangular; posterior 1/2 of segment expanded dorsally into rectangular plate; each anterodorsal margin deeply concave in lateral view; each ventral margin uniformly convex, without incision at vertical apodeme; each posterior margin nearly straight, starting below cercus; ventral side of posterior-most part of segment IX with setae ( Fig. 152 ). In dorsal view segment IX with narrow anterior lobes; in dorsal view anterodorsal margin forming deep, wide, U-shaped incision with anterad-orienting processes on each side. In ventral view segment IX nearly quadrangular, with concave anterior margin and straight posterior margin; posterior margin without central projection. Tergum X simple; apical 1/3rd much more slender than basal 2/3rds ( Fig. 150 ); tergum X orienting posterad, slightly curved ventrad along its length; divided into 2 lateral branches from base in dorsal and ventral view ( Figs. 151, 152 ); each branch uniformly narrow in dorsal and ventral view, except apical 1/3rd narrower; without dorsal process in lateral view, but with narrow, hook-shape lateral process on each lateral branch at distal 1/3rd; pair of sensillae located on dorsal end of thickest part of lateral process. Cerci nearly club-shaped in lateral view, located at 2/3rds height of posterior margin of segment IX and oriented dorsad in lateral view and posterad in dorsal view; covered by long setae. Gonopods as long as segment IX, 3-branched. Each dorsal branch long, slightly exceeding tergum X in lateral view; uniformly slender, bending posterad at mid-length; anterodorsal margin convex; ventral margin slightly concave; in ventral view slightly curved mesad; without mesal megasetae. Ventral branch of each gonopod nearly rectangular in lateral view, with undulating posterior margin and smooth ventral margin; mesal margins strongly convex in ventral view. Mesal branches darkly pigmented, slightly longer than each circus; straight in lateral view; curved mesad in ventral view. Phallic apparatus slightly longer than rest of genitalia: phallotheca, in lateral view, with anterior part 2x thicker than posterior part; in ventral view anterior part about 1.5x wider than posterior part; apicoventral spine absent on phallotheca; large phallotremal sclerite in phallic apparatus forming complex structure in lateral view; in ventral view forming paired rays separated by narrow plate; 2 short, nearly black endothecal spines present.

Female. Unknown.

Holotype male: VITI LEVU: Vuda Prov., Koroyanitu N.N.P. Abaca Village, Malaise trap, 6–26.v.2003, 17°40’S, 177°33’E [17.3333°S, 177.5500°E], 400 m, leg. M. Irwin, E. Schlinger & M. Tokota’a [loc#01] [ FNIC].

Paratypes: VANUA LEVU : Bua Prov., Kilaka, Malaise trap, 24.vi–21.vii.2004, 16°48’927’’S, 178°59’110’’E [16.8258°S, 178.9864°E], 146 m, M. Irwin, E. Schlinger & M. Tokota’a [loc#24] — 1 male [ NHRS, DNA voucher IK9]. TAVEUNI: Cakadrove Prov., 5.6 km SE Tavuki Village, Malaise trap in rainforest, 31.x–14.xi.2002, 16.843°S, 179.955°W, 1187 m, E. Schlinger & M. Tokota’a [loc#39] — 1 male [ BPBM].

Etymology: Tipulida , named after the Diptera family Tipulidae (crane flies) into which the genus Trichoptera Strobl was described.

Distribution: Taveuni, Viti Levu and Vanua Levu.

Figures

FIGURES 22 – 29. Right wings, dorsal views. 22 — C. cartwrighti, new species, holotype; 23 — C. kimminsi, new species, holotype; 24 — C. kadavuensis, new species, holotype; 25 — C. lavensis, new species, holotype; 26 — C. devoensis, new species, holotype; 27 — C. helomyzida, new species, holotype; 28 — C. tipulida, new species, holotype; 29 — C. psychodida, new species, holotype.

FIGURES 150 – 154. Chimarra tipulida, new species, holotype. 150 — genitalia, left lateral view; 151 — genitalia, dorsal view; 152 — genitalia, ventral view; 153 — phallic apparatus, left lateral view; 154 — phallic apparatus, ventral view.