Clivina saundersi Andrewes, 1926

Balkenohl, Michael, 2021, Belgian Journal of Entomology, Belgian Journal of Entomology 115, pp. 1-83 : 26-28

Clivina saundersi Andrewes, 1926

( Figs 7 , 14 , 21 , 41 , 61 , 88 )

Clivina saundersi ANDREWES, 1926 C: 273, 1930: 116; CSIKI, 1927: 553; LORENZ, 1998: 133, 2005: 144; BALKENOHL, 2001: 16.

TYPE MATERIAL. Neotype by present designation: • ♂, with labels and data: white, black printed “ Tjinta Radja ” / white, black printed “ Sumatra: Mjöberg ” / white, handwritten with black ink and black printed “ Saundersi Cotype Andr H.E. Andrewes det.” / blue, black printed “4237 E91 +” / white, black printed “ NHRS- JLKB 000065315 ” ( Fig. 14 ) ( NHRS) .

REMARKS. The specimen shows a pin hole on the right elytron but is complete.

ANDREWES (1926c) based his description on three specimens. The holotype was collected in Singapore by C. J. Saunders and was part of Andrewes’ collection, now deposited in BMNH. While on research loan in 2008, it was in a bag that was stolen from the borrower at a meeting of Entomologists in Prague, and is lost (personal communication from Maxwell V. L. Barclay, Head of Coleoptera at the BMNH). The ‘cotype’ ( ANDREWES, 1926c) from Kuala Lumpur, Bukit Cherakah, was deposited in the Federal Malay States Museum and is not available. So, the ‘cotype’ collected by E. Mjöberg in Sumatra, Tjinta Radja, deposited in NHRS is the only available ‘cotype’. It fits exactly with the description of ANDREWES (1926c) and is here designated as Neotype with the express purpose of clarifying the taxonomic status and the type locality of Clivina saundersi . Iam satisfied that this designation meets the conditions of ICZN Article 75, and as a surviving paratype, it also meets the specifications of ICZN Recommendation 75A ( ICZN, 1999).

DIAGNOSIS. Amedium sized species with subcrenulated lateral margin of the pronotum and crenulated margin of the elytron in anterior quarter, and isodiametric reticulation on all intervals of the elytron. Distinguished from all other species by the tubercles at the base of intervals two, three and four, by the first setigerous puncture on the elytron which is situated at the third stria, and by the interval eight which is carinate for its whole length.

REDESCRIPTION. Measurements. Lectotype: Body length 8.87 mm, width 2.77 mm; ratio length/widthofpronotum 0.91; ratiolength/widthofelytra 1.71.

Colour: Head semiopaque, pronotum glossy, elytron opaque. Piceous. Intermediate and hind leg dark fuscous, antenna and tarsomeres fuscous.

Head: More than two-thirds width of pronotum. Clypeus slightly bilobed anteriorly, with obtuse notch at middle and lateral angle rounded; wing less protruding as clypeus, somewhat elongated, slightly hollowed out concavely, separated from clypeus by distinct notch; supraantennal plate convex; clypeus, wing and supraantennal plate slightly reflexed margined. Supraantennal plate extended up to mid-eye level, with rugae at mid-eye level, separated from wing by sharp notch. Clypeus square, with transverse elevation, separated from frons by distinct furrow. Frons moderately convex, with longitudinal pore at middle, with transverse flat impression, with two parallel running blunt lateral carinae at posterior eye-level. Clypeus and frons with some fine rugae and scattered punctures, separated from wing and supraantennal plate by deep and conspicuously wide furrow continuing posteriorly as wide supraorbitalfurrow with two supraorbital setigerous punctures at posterior eye level. Neck constriction developed as distinct step, with band of punctures. Eye laterally projecting. Gena distinct. Antenna reaching posterior angle of pronotum, antennomeres four to ten elongate (ratio L/ W 2.03). Labrum convex anteriorly, with isodiametric reticulation, seven setose. Mentum with indistinct longitudinally reticulated surface, with lobe elongated and slightly hollowed out, shape of lobe like a skewed trapezium with rounded antero-lateral angle, reflexed margined anteriorly, carinate laterally, lateral carina somewhat removed from margin, with sharp carina at middle, median tooth wide, distinctly less protruding anteriorly as lobe, obtusely angled anteriorly.

Pronotum ( Fig. 61 ): Disk distinctly convex in lateral and frontal view. Wider than long. Reflexed lateral margin subcrenulated, slightly convex, conspicuously attenuated in anterior two thirds, widest at posterior third; anterior angle obtusely rounded off, slightly projecting; posterior angle developed as distinct somewhat rounded tooth. Lateral channel with series of pits between the two setigerous punctures, slightly widened between anterior and posterior setigerous puncture. Median line narrow, line-like; anterior transverse line wider, joining median line, joining basal constriction. Surface with transverse wrinkles, with minute scattered punctures, base granulated at declivity.

Elytron: Disk flattened in anterior half in lateral view, convex on frontal view. Less than twice as long as wide, not dilated in apical half, maximum width at middle. Reflexed lateral margin with few small notches in anterior quarter, smooth posteriorly. Scutellar striole distinct. Base of elytron with setigerous tubercle at base of first stria, with small tubercle at base of second interval, with distinct tubercle at base of interval three and four, each. Striae moderately deep, punctuate-striate, one to three free at base, four to seven joining at humerus, one joining with seven at apex, two to six ending free at apex. Intervals moderately convex, seven with short carina at humerus, eight carinate in its whole length. Third interval with four setigerous punctures, all approaching third stria. Surface of all intervals semiopaque, covered with faint isodiametric reticulation.

Hind wings: Fully developed.

Lower surface: Proepisternum covered completely with fine but distinct isodiametric reticulation and laterally with transverse wrinkles. Terminal sternite of abdomen developed apically as deep narrow transverse fold.

Legs: Legs covered with isodiametric reticulation. Protibia with three slender spines of moderate length, sulcate dorsally, movable spur regularly arcuate.

Male genitalia ( Fig. 88 ): Median lobe of moderate length, moderately broad, regularly arcuate in basal two thirds, straight apically, ventral rim distinct from basal quarter to apical third, apex duckbill like developed, slightly wider apically. Endophallus posteriorly with bristles. Ventral paramere moderately wide, lateral apophysis small, the other one elongated, apex regularly tapered, with three setae at apex of decreasing length, with one small seta preapically. Dorsal paramere regularly arcuate, with slender apophyses, at apex with four setae of different length.

Female coxostylus and laterotergite: Unknown.

VARIATION. Unknown.

DISTRIBUTION. Known from the type locality Tjinta Radja in Sumatra. The lost holotype was found in Singapore.

Figures

Fig. 7. Clivina saundersi Andrewes, 1926. Base of the elytra in dorsal aspect. White arrow: setigerous tubercle, yellow arrows: additional tubercles situated on top of the base and slightly down the declivity.

Figs 9–15. Clivina species, labels of primary historic type material. 9. C. castanea Westwood, 1837, holotype; 10. C. clivinoides Schmidt-Göbel, 1846, lectotype; 11. C. parryi Putzeys, 1863, lectotype; 12. C. lata Putzeys, 1866, lectotype; 13. C. agona Putzeys, 1866, holotype; 14. C. saundersi Andrewes, 1926, neotype; 15. C. trachys Andrewes, 1930, holotype.

Fig. 21. C. saundersi Andrewes, 1926, neotype.

Fig. 41. C. saundersi Andrewes, 1926, neotype.

Fig. 61. C. saundersi Andrewes, 1926, neotype.

Fig. 88. C. saundersi Andrewes, 1926, neotype.