Eocypraea undetermined, (LAMARCK, 1802)

Groves, Lindsey & Squires, Richard L., 2023, Revision of northeast Pacific Paleogene cypraeoidean gastropods, including recognition of three new species: implications for paleobiogeographic distribution and faunal turnover, PaleoBios 40 (10), pp. 1-52 : 27

EOCYPRAEA SP. CF. E. INFLATA ( LAMARCK, 1802)

FIGS. 8G–I

Eocypraea castacensis ( Stewart, 1926 [1927]). Vokes, 1939, p. 154 [in part], pl. 20, fig. 14 [not fig. 9, = Bernaya kaylinae n. sp.].

Cypraea castacensis (Stewart) . Ingram, 1942. p. 103 [in part], pl. 8, fig. 6 [not fig. 5, = Bernaya kaylinae n. sp.].

Eocypraea (Eocypraea) moumieti Dolin and Dolin, 1983 . p. 36. Groves, 1997. p. 8. Not Eocypraea moumieti Dolin and Dolin, 1983 . Not Eocypraea inflata var. moumieti Dolin and Dolin, 1983 . p. 46.

Eocypraea (Eocypraea) sp. cf. E. (E.) inflata ( Lamarck, 1802) . Groves, 2011. p. 50, figs. 11, 12. Groves and Squires, 2021. p. 231.

Referred Specimen—Hypotype (of Vokes, 1939, of Ingram, 1942, of Groves, 2011) UCMP 15815 ( Figs. 8G–I ), length 27.3, width 18.7 mm, height 15.1, UCMP Locality A-1282.

Occurrence— Domengine Formation, middle Eocene, Reef Ridge, Fresno County, central California .

Etymology— Originally from Latin inflata for the highly inflated shell.

Remarks— The hypotype of Vokes (1939), Ingram (1942), and Groves (2011) most closely resembles Eocypraea inflata ( Lamarck, 1802) from middle Eocene (Lutetion/Bartonian) strata of France, Belgium, and England. This is particularly evident from the illustrations of Cossmann (1903: pl. 9, figs. 18, 19), Cossmann and Pissaro (1911: pl. 32, fig. 162-7), and Wenz (1941: fig. 2882). It superficially resembles Luponovula maniobraensis ( Squires and Advocate, 1986) from the lower Eocene (“Capay Stage”) Maniobra Formation of Riverside County, southern California. Luponovula maniobraensis is more elongate, has coarser dentition, has a prominent basal callus, and is larger than Eocypraea sp. cf. E. inflata .

Dolin and Dolin (1983: p. 37, table on p. 46 [as Eocypraea inflata var. moumieti ]) reported Eocypraea moumieti Dolin and Dolin, 1983 , which is known from the Gan Basin, Pyrénées Atlantique Department, France, as also occurring in California. They reported that this California occurrence is questionably “ castacensis ” Vokes, 1939, but Vokes was not the author of castacensis . That particular specimen (hypotype UCMP 33808 of Vokes, 1939) is now Bernaya kaylinae n. sp. In summary, E. sp., cf. E. inflata ( Lamarck, 1802) occurs very rarely in the NEP region, whereas E. moumieti Dolin and Dolin, 1983 does not occur in this region.

Perrilliat et al. (2003: p. 48, pl. 2, figs. 33, 34) reported a specimen of a middle Eocene cypraeoidean as E. (E.) inflata ( Lamarck, 1802) [hypotype IHN 6736] from the San Juan Formation, Chiapas, southern México. Unfortunately, this specimen is too poorly preserved to afford a positive identification.

Figures

Figure 8A–C. Eocypraea novasumma (Nelson, 1925), Santa Susana Formation (Paleocene, Thanetian Stage), Runkle Canyon, north side Simi Hills, Ventura County, southern California. Basal (A), dorsal (B), and right-labral (C) views, 3.2x, length 17 mm, width 11.6 mm, height 9.3 mm, of holotype UCMP 30499, UCMP Locality 4776. D–F. Eocypraea takeosusukii Groves, 2011, “Martinez” Formation (Paleocene, Thanetian), Lower Lake area, Lake County, northern California. Basal (D), dorsal (E), and left-lateral (columellar) views, 2.6x, length 18.8 mm, width 14.4 mm, height 11.2 mm, of holotype LACMIP 7045.1, LACMIP Type 13645, LACMIP Locality 7045 [ex CIT Locality 1580]. G–I. Eocypraea sp., cf. E. inflata (Lamarck, 1802), “Domengine Formation” (Eocene, “Domen- gine Stage”), Reef Ridge, Kings County, central California. Basal (G), dorsal (H), and right (labral) (I) views, 1.8x, length 26 mm, width 17.7 mm, height 13 mm, of hypotype UCMP 15815, Locality UCMP A-1282. J–L. Eocypraea sp. Lodo Formation (basal part) (Paleocene, Thanetian Stage), Tumey Hills, Fresno County, central California. Basal (J), dorsal (K), and right (labral) (L) views, 1.5x, length 33 mm, width 21.5 mm, height 19 mm, of hypotype CASG 61666.04 [ex LSJU 10236], CASG Locality 61666 [ex LSJU Locality 2073; = UCMP Locality A-9717]. M–P. Grovesia castacensis (Stewart, 1926 [1927]), Tejon Formation (Eocene, “Tejon Stage”), Tehachapi Mountains, Kern County, southern California. Basal (M), dorsal (N), and left (columellar) (O) views, 4.5x, length 12.2 mm, width 7.8 mm, height 5.9 mm, of holotype UCMP 11690, Grapevine Canyon, UCMP Locality 452. Basal (P) view, 3.4x, length 15.5 mm, width 9.1 mm, height 6.9 mm, of hypotype LACMIP 41206.2, LACMIP Type 14941, base of Metralla Sandstone Member, Live Oak Canyon, LACMIP Locality 41206.