Togoda Melichar, 1906

Gnezdilov, Vladimir M., 2025, Review of the genus Togoda Melichar, 1906 (Hemiptera, Auchenorrhyncha, Fulgoroidea, Tropiduchidae), with descriptions of three new species from tropical Africa and Madagascar, Zoosystema 47 (23), pp. 473-488 : 474

Genus Togoda Melichar, 1906

Togoda Melichar, 1906: 233 .

TYPE SPECIES. — Togoda africana Melichar, 1906 , by monotypy.

DISTRIBUTION. — Tropical Africa ( Nigeria, Ghana, Togo, Central African Republic, South African Republic, Mozambique, Tanzania: Zanzibar Island) and Madagascar ( Fig. 11 ).

COMPOSITION. — Five species.

DIAGNOSIS. — Coryphe transverse. Metope long and wide, nearly parallel-sided (in dorsal and frontal views), protruding in shape of wide proboscis (in lateral view). Coryphe and metope joint at obtuse angle in lateral view. Postclypeus large, with median carina. Metopoclypeal suture straight. Scapus short, cylindrical. Pedicel 4-5 times longer than scapus, elongate, twice as long as wide, slightly convex dorsally, covered with setae and sensory organs ( Fig. 1D ). Ocelli absent. Rostrum long, narrow, reaching hind coxae or slightly surpassing them; third segment slightly longer than second one, narrowing and bent apically ( Fig. 7B ). Pronotum large, with distinct lateral carinae diverging posteriorly, wide paradiscal fields behind the eyes and wide paranotal lobes; posterior margin strongly concave. Median part of pronotum hanging over coryphe. Mesonotum 1.5 times as long as pronotum, without carinae. Forewing with loop-shaped process of precostal margin basally and with semicircular lobe on wing margin behind claw-shaped apex of clavus, both with marginal setae ( Figs 1 ; 2 ). Marginal cell below the semicircular lobe distinctly concave. Tegulae large. Fore wings without hypocostal plate, but with narrow precostal field, with transverse veins and with a wide subcostal field in its basal fifth; clavus open. Hind wings well developed, slightly shorter than fore wings ( Fig. 7A ). Hind tibia with two large lateral spines on the apical half and one small lateral spine on the proximal half. First and second metatarsomeres nearly equal in length, with dense long hair-shaped setae ventrally. Second metatarsomere with only two latero-apical spines. Pygofer with widely and slightly concave lower margin ( Fig. 9J ). Penis strongly curved in lateral view. Apical part of aedeagus weakly sclerotized and apparently sometimes swollen. Styles massive, fused at its basal halves ( Fig. 9I ), with short capitulum, without neck and with a strong comb on the margin below capitulum. Finger-shaped process of the style large, curved and turned posteriorly and ventrally. Abdominal sternite III large, with triangularly protruding hind margin overhanging above other sternites ( Figs 4B, D ; 5B, D ). Female sternite VII with weakly convex medially hind margin. Gonoplacs large, triangular-shaped, flat, with large marginal teeth ( Fig. 3C, D ).

Figures

FIG. 11. — Distribution of the genus Togoda Melichar, 1906. Abbreviations: �, Togoda sp.; �, T. africana Melichar, 1906; �, T. carinata n. sp.; �, T. excisa n. sp.; �, T. minima n. sp.; �, T. transvaalensis Schmidt, 1910. Not to scale.

FIG. 1. — Forewing and pedicel structure of Trienopini: A, Trienopa flavida Signoret, 1860 (Madagascar), male, basal part of forewing, with ear-shaped process of precostal margin; B, Togoda sp. (Ghana), female, pronotum and basal part of forewing, with ear-shaped process of precostal margin; C, Togoda sp. (Ghana), female, ear-shaped process of precostal margin dorsally; D, Togoda sp. (Ghana), female, pedicel; E, Togoda sp. (Ghana), female, precostal margin of forewing at its middle; F, Togoda sp. (Ghana), female, ear-shaped process margin. Scale bars: A, 400 μm; B, 500 μm; C, 200 μm; D, E, 100 μm; F, 50 μm.

FIG. 7. — Togoda africana Melichar, 1906, Togo (Bismarckburg), male: A, right hind wing; B, rostrum. Abbreviations: A1, first anal vein; A2, second anal vein; CuA, cubitus anterior; CuP, cubitus posterior; M, median vein; Pcu, postcubitus; R, radial vein. Scale bars: A, 1 mm; B, 0.5 mm.

FIG. 2. — Forewing structure of Togoda spp.: A, B, Togoda africana Melichar, 1906, Togo (Bismarckburg), male: A, apex of forewing clavus, dorsal view; B, semicircular process of forewing margin behind claval apex; C, D, Togoda excisa n. sp., female paratype: C, semicircular process of forewing margin behind claval apex; D, margin of semicircular process. Scale bars: A, 500 μm; B, 100 μm; C, 400 μm; D, 50 μm.

FIG. 9. — Togoda excisa n. sp., holotype, male genitalia: A, pygofer and anal tube, lateral view; B, anal tube, lateral view; C, anal tube, dorsal view; D, penis, left lateral view; E, penis, ventral view; F, penis, right lateral view; G, style, lateral view; H, capitulum of style, dorsal view; I, styles, ventral view; J, pygofer, caudal view. Abbreviations: at, apical tooth of style; b, c, processes of aedeagus; lt, lateral tooth of style. Not to scale.

FIG. 4. — Habitus of Togoda spp.: A, B, Togoda africana Melichar, 1906, Togo (Bismarckburg), male, dorsal view (A) and lateral view (B); C, D, Togoda excisa n. sp., female paratype, dorsal view (C) and lateral view (D). Scale bar: 1 mm.

FIG. 5. — Habitus of Togoda spp.: A, B, Togoda minima n. sp., female paratype, dorsal view (A) and lateral view (B); C, D, T. carinata n. sp., holotype, dorsal view (C) and lateral view (D). Scale bar: 1 mm.

FIG. 3. — Habitus of Trienopini:A, Trienopa typica (Distant, 1909) (Seychelles); B, Togoda sp. (Nigeria:Ekiti, Ikere); C, Togoda sp. (Ghana), ovipositor, lateral view; D, Togoda sp. (Ghana), female, abdomen, ventral view. Scale bars: 1 mm. Credits: A, Gernot Kunz; B, Babajide Agboola.