Togoda carinata, Gnezdilov, 2025

Gnezdilov, Vladimir M., 2025, Review of the genus Togoda Melichar, 1906 (Hemiptera, Auchenorrhyncha, Fulgoroidea, Tropiduchidae), with descriptions of three new species from tropical Africa and Madagascar, Zoosystema 47 (23), pp. 473-488 : 482-487

Togoda carinata n. sp.

( Figs 5C, D ; 6H, G )

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TYPE MATERIAL. — Holotype. Madagascar • ♀; “Madagascar Est / distr. Mananara – N. / Antanambe VII / Vadon et Peyrieras” [print with hand written date] // “Museum Paris / MNHN(EH): / 16570” [print]; MNHN.

DIAGNOSIS. — Metope with two weak carinae medially and groove between them. Postclypeus with strong triangular-shaped median carina. Fore and middle femora flattened or nearly foliate.

ETYMOLOGY. — Species name referring to strong carina of postclypeus.

DISTRIBUTION. — Northeastern Madagascar (Mananara Nord District).

DESCRIPTION

Structure ( Figs 5C, D ; 6H, G )

Coryphe three times as wide as long medially, smooth, without carinae; anterior margin convex; posterior margin concave ( Fig. 5C ). Metope with two weak carinae medially and groove between them, with sparse pustules ( Fig. 6G ). Coryphe-metope margin keel-shaped. Postclypeus with strong triangular-shaped median carina ( Fig. 5D ; 6H ). Anteclypeus with fine median carina. Eyes large, diameter of each eye 1.5-2 times as width of coryphe. Fore wings nearly oval, protruding beyond ovipositor apex ( Fig. 5D ). Basal cell small and oval. Radius and median running from the same point on the basal cell. Radius and median firstly furcating closely to basal cell in basal wing fifth. Many transverse veins between branches of median vein. Forewing vein branching sequence: R 5; r-m 6; M 5; CuA 2, furcating near to wing middle. Fore and middle femora flattened or nearly foliate. Hind tibia with ten spines apically. First metatarsomere with two latero-apical and five intermediate spines arranged in arc apically.

Coloration ( Fig. 5C, D )

Generally brown to dark brown. Coryphe reddish brown. Metope dark brown, with yellowish pustules. Head laterally with wide black stripe above antennae. Scapi, pedicels, and postclypeus dark brown to black. Anteclypeus light yellow. Rostrum light yellow, with black apex. Pro- and mesonotum reddish brown to dark brown. Fore wings reddish brown, with large oval black spot on each clavus, between Pcu and first anal vein, and with several large black spots inside of cells on corium including oval one between posterior branch of median and CuA in front of claval spot. Hind wings dark brown. Fore and middle coxae, femora, and tibiae dark brown. Trochanters light yellow. Hind femora dark brown. Hind tibiae dark brown proximally and light yellow distally. First and second tarsomeres of fore and middle tarsi light yellow; third tarsomeres dark brown. Hind tarsi light yellow. Claws dark brown to black. Apices of leg spines black. Abdominal sternites dark brown. Ovipositor yellowish brown.

Measurement

Total length. Female: 5.5 mm.

Figures

FIG. 5. — Habitus of Togoda spp.: A, B, Togoda minima n. sp., female paratype, dorsal view (A) and lateral view (B); C, D, T. carinata n. sp., holotype, dorsal view (C) and lateral view (D). Scale bar: 1 mm.

FIG. 6. — Head of Togoda spp.: A, B, T. africana Melichar, 1906, male, Togo (Bismarckburg), head, lateral view (A), metopial proboscis, frontal view (B); C, D, T. excisa n. sp., paratype, head, lateral view (C), metopial proboscis, frontal view (D); E, F, T. minima n. sp., paratype, head, lateral view (E); metopial proboscis, frontal view (F); H, G, T. carinata n. sp., metopial proboscis, frontal view (G); holotype, head, lateral view (H). Scale bar: 0.5 mm.