Togoda minima, Gnezdilov, 2025

Gnezdilov, Vladimir M., 2025, Review of the genus Togoda Melichar, 1906 (Hemiptera, Auchenorrhyncha, Fulgoroidea, Tropiduchidae), with descriptions of three new species from tropical Africa and Madagascar, Zoosystema 47 (23), pp. 473-488 : 480-482

publication ID	https://doi.org/10.5252/zoosystema2025v47a23
publication LSID	lsid:zoobank.org:pub:44B9F8A0-391B-4C00-AE1B-23D3E5494409
DOI	https://doi.org/10.5281/zenodo.17259625
persistent identifier	https://treatment.plazi.org/id/911087DF-1211-FFB3-3B0F-FB6A217CF864
treatment provided by	Plazi
scientific name	Togoda minima
status	sp. nov.

Treatment

Togoda minima n. sp.

( Figs 5A, B View FIG ; 6E, F View FIG ; 10 View FIG )

urn:lsid:zoobank.org:act:

TYPE MATERIAL. — Holotype. Mozambique • ♂; “Museum Paris / Moçambique / Vallée du Pungouí / Guengére / G. Vasse 1906” [grey blue] // “Février” [white] // “Museum Paris / MNHN (EH) / 16572” [white]; MNHN.

Paratype. Mozambique • ♀; “Museum Paris / Mozambique / Vila Pery / P. Lesne 1928” [grey blue] // “Mars” [white] // “Museum Paris / MNHN (EH) / 16571” [white]; MNHN .

DIAGNOSIS. — Fore wings small, with 1/5 of its length behind apex of clavus in lateral view. Metopial proboscis without notch frontally. Phallobase long, nearly as long as aedeagus, asymmetric.

ETYMOLOGY. — Species name referring to small fore wings.

DISTRIBUTION. — Central Mozambique (Nhamatanda District).

DESCRIPTION

Structure ( Figs 5A, B View FIG ; 6E, F View FIG )

Coryphe weakly transverse, 1.5 times as wide as long medially, without carinae, anterior margin straight, posterior margin nearly straight ( Fig. 5A View FIG ). Metope, pro- and mesonotum densely covered with pustules ( Fig. 6F View FIG ). Coryphe-metope margin keel-shaped ( Figs 5B View FIG ; 6E View FIG ). Metopial proboscis without notch frontally ( Fig. 6F View FIG ). Post- and anteclypeus with fine median carina medially which is running from the middle of postclypeus throughout anteclypeus. Eyes large, diameter of each eye nearly as wide as coryphe. Median part of pronotum with a groove medially. Fore wings rather short, nearly oval, surpassing by 1/5 of its length beyond abdominal apex in male and reaching abdominal apex in female ( Fig. 5B View FIG ). Basal cell narrowly oval. Radius and median running from basal cell by one short stem or from one point and firstly furcating near to the cell in basal fifth of wing. Forewing vein branching sequence: R 5, all branches terminating at costa not reaching wing margin; r-m 4; M 10, firstly furcating near to basal cell, M 1 and M 2 again furcating after wing middle; im 12; m-cua 4-6; CuA 2, furcating near wing middle; icua 3. Pcu joint A 1 near to wing middle. Clavus long, 4/5 of wing length. Legs not flattened neither foliated. Hind tibia with nine spines apically. First metatarsomere with two latero-apical and 7-9 intermediate spines arranged in an arc apically.

Coloration ( Fig. 5A, B View FIG )

General coloration yellowish brown. Head laterally light yellow below antennae, with wide black stripe above scapi. Coryphe, pronotum, and mesonotum yellowish brown. Each paranotal lobe of pronotum with large dark brown to black area proximally. Mesonotum sometimes with dark brown lateral parts. Metope brown to dark brown, with reddish lateral margins and light yellow pustules. Postclypeus yellowish brown to dark brown, with pale median carina. Anteclypeus light yellow. Rostrum light yellow, with black apex. Scapi and pedicels brown. Each paranotal lobe of pronotum with large dark brown to black area proximally. Mesonotum sometimes with dark brown lateral parts. Forewing yellowish brown to dark brown, with basal subcostal field brown to dark brown, some cells of corium sometimes dark brown to black, and veins reddish brown. Basal processes of forewing precostal margins dark brown to black. Hind wings dark brown. Fore femora light yellowish brown to dark brown. Fore tibiae with three dark brown bands. Fore tarsi dark brown. Apices of leg spines black. Claws dark brown. Thorax and abdomen below light yellow. Abdominal sternite III light yellow. Abdominal sternites IV-VI yellowish brown. Female sternite VII with black laterally hind margin. Gonoplac margins dark brown, with black teeth. Gonocoxae VIII dark brown, with black hind margins proximally. Male genital block yellowish brown.

Male terminalia ( Fig. 10 View FIG )

Pygofer wide in dorsal half, with strongly convex hind margins (in lateral view) ( Fig. 10A View FIG ). Anal tube elongate, twice as long as wide medially, narrowing to truncate apex (in dorsal view) ( Fig. 10C View FIG ).

Apex of anal tube slightly turned downwards (in lateral view) ( Fig. 10B View FIG ). Anal column short and wide. Penis wide, horse-shoe shaped (in lateral view) ( Fig. 10D, E View FIG ). Phallobase long, nearly as long as aedeagus, assymetric, with ventral lobe shifted to the left side and narrowing apically. Aedeagus with a single hookshaped subapical process turned downwards ( Fig. 10D, E, a View FIG ) on right side. Phallobase with long and wide plate-shaped process, with a short tooth-shaped projection apically, turned upwards on right side ( Fig. 10D, e View FIG ) and narrow pointed apically and turned upwards process on left side ( Fig. 10D, E, f View FIG ). Style with nearly straight hind margin and large apical tooth ( Fig. 10F, G View FIG , at). Connective with elongate large cup ( Fig. 10D View FIG ).