Anthemis tinctoria subsp. fussii

Tutin, T. G., Heywood, V. H., Burges, N. A., Moore, D. M., Valentine, D. H., Walters, S. M. & Webb, D. A., 1976, Flora Europaea. Volume 4. Plantaginaceae to Compositae (and Rubiaceae), Cambridge University Press : 156

(d) Subsp. fussii (Griseb.) Beidie, FI. Veg. Muni. Bucegi 270 (1967)

( A. tinctoria var. fussii Griseb. ):

Like subsp. (a) but the involucral bracts all acute or subacute with a distinct dark-brown or blackish margin and apex; outer receptacular scales suffused with brown towards the apex; corona longer. • S. Carpathians.

Somewhat intermediate between 42(a) and 44 and perhaps a local variant of the latter.

Plants from E.C. Europe have leaves with long, narrow segments and sometimes also dense greyish indumentum and pale yellow ligules (at least in the dry state) as in subsp. (b).

A. markhotensis Fedorov in Schischkin & Bobrov, Fl. URSS 26: 867 (1961) and A. zephyrovii Dobrocz. , Ukr. Bot. Žur. 18(2): 70 (1961) are taxa of uncertain status closely related to 42(b). The former has leaves 3-4 cm, with wide segments and flat lobes and capitula 20-25(-30) mm in diameter with golden-yellow ligules, and the latter has leaves of similar size but crowded near the middle of the stem, and capitula 15—17(—25) mm in diameter with pale yellow ligules. Both occur in Krym.