Galium papillosum, Lapeyr.

Tutin, T. G., Heywood, V. H., Burges, N. A., Moore, D. M., Valentine, D. H., Walters, S. M. & Webb, D. A., 1976, Flora Europaea. Volume 4. Plantaginaceae to Compositae (and Rubiaceae), Cambridge University Press : 30-31

97. G. papillosum Lapeyr. , Hist. Abr. Pyr. 66 (1813).

Like 94 but dark green to brownish when dry, with short, subterranean stolons; stems (10-)20-60(-80) cm, more than 1 mm in diameter, more or less retrorsely aculeolate, but also hairyin places, or sometimes entirely glabrous and smooth; middle internodes 2-4 times as long as the leaves; leaves (8—) 10—25(—30) x (0-8-) l-2-2(2-8) mm, in whorls of (7-)8-9(-10), 9—14 times as long as wide, thin to somewhat coriaceous, upper surface and margin antrorsely scabrid or hairy; inflorescence broadly pyramidal, branches from lower third of stem; pedicels 0-5-l-2 mm, flowers crowded; corolla 1 -5-2-5 mm in diameter; fruit c. 1 -5 mm, smooth to obtusely papillose, more or less shining. In = 22,44. Dry places. • N.E. Spain, E. Pyrenees. Ga Hs.

Octoploid individuals (2л = 88) with stems not retrorsely aculeolate form transitions to 99, especially in the mountains and on the N. side of the Pyrenees.