Leontodon crispus, Vili.

Tutin, T. G., Heywood, V. H., Burges, N. A., Moore, D. M., Valentine, D. H., Walters, S. M. & Webb, D. A., 1976, Flora Europaea. Volume 4. Plantaginaceae to Compositae (and Rubiaceae), Cambridge University Press : 314

19. L. crispus Vili. , Prosp. Pl. Dauph. 34 (1779).

Perennial with branched, oblique stock and long, vertical rhizome. Stems 1-6,7-40 cm, more or less thickened at apex, simple or branched, with few to numerous, rigid, stalked 2- to 4-fid hairs (often with only their bases remaining) and sometimes shorter, weaker, simple eglandular and sessile stellate hairs; bracts 0-5. Leaves numerous, 20-140 x 3-15(-20) mm, more or less oblanceolate, dentate to pinnatifid, usually with numerous 2- to 7-fid hairs, sometimes also with sessile stellate hairs. Capitula l (-3). Involucre 10-25 x 7-14 mm; bracts oblong-lanceolate, more or less acute, glabrous or with simple eglandular, 2- to 4-fid or stellate hairs, often pectinate-ciliate. Ligules yellow, sometimes reddish-purple on outer face. Stigmas yellow. Achenes 7-12 mm, brown, with beak up to about as long as remainder of achene, with short rigid hairs above and transversely muricate below; pappus of 2 rows of very pale brown plumose hairs dilated at the base, the outer shorter than the inner. S'. & S.E. Europe, extending northwards to c. 55° N. in E.C. Russia. Al Bu Co Ga Gr Hs It Ju Rm Rs (C, W, K, E) Si Tu.

1 Leaves with at least some sessile stellate hairs and numerous 2- to 7-fid stalked hairs 2 Leaves withscatteredsmallstellatehairs andnumerous stalked 2- to 5-fid hairs (d) subsp. bourgaeanus

2 Leaves with numerous or dense ± sessile stellate hairs and 3-

to 7-fid hairs (e) subsp. graecus 1 Leaves with stalked 2- to 7-fid hairs, without sessile stellate hairs

3 Involucral bracts with minute hairs or glabrous

(a) subsp. rossianus 3 Involucralbractswithrigidhairs,at leastonthemargins

4 Involucre 12-15 mm, the bracts not or minutely pectinate-ciliate; achenes 7-12 mm (b) subsp. crispus 4 Involucre 14-25 mm, with at least the outer bracts strongly pectinate-ciliate; achenes 12-20 mm (c) subsp. asperrimus