Centaurea margaritacea subsp. protomargaritacea

Tutin, T. G., Heywood, V. H., Burges, N. A., Moore, D. M., Valentine, D. H., Walters, S. M. & Webb, D. A., 1976, Flora Europaea. Volume 4. Plantaginaceae to Compositae (and Rubiaceae), Cambridge University Press : 286

(e) Subsp. protomargaritacea (Klokov) Dostál, loc. cit. (1976)

( C. protomargaritacea Klokov ):

Stems smooth above. Leafsegments linear-filiform. Involucre 11-16 x 11-15 mm, ovoid; appendages 5-6 x 4-5 mm, not completely covering bracts, elliptic-orbicular, yellow or pale orange, membranous, not pellucid, the centre brown, convex, the margin lacerate, convolute, the apex muticous. Florets pink, to almost white. Achenes 3-4 mm, blackish, with white ribs; pappus c. 3 mm, somewhat shorter than achene. Submaritime sands. • S. Ukraine ( E. side of estuary of R. Bug).