Centaurea margaritacea subsp. appendicata

Tutin, T. G., Heywood, V. H., Burges, N. A., Moore, D. M., Valentine, D. H., Walters, S. M. & Webb, D. A., 1976, Flora Europaea. Volume 4. Plantaginaceae to Compositae (and Rubiaceae), Cambridge University Press : 286

(f) Subsp. appendicata (Klokov) Dostál, loc. cit. (1976)

( C. appendicata Klokov ):

Stems scabrid above. Leaf-segments broadly linear. Involucre 22-27 mm in diameter, globose; appendages 10-12 x 6-11 mm, not covering bracts, orbicular- rhombic, yellow, keeled, coriaceous on keel, with oblongtriangular, dark purple central spot, the margin wide, pellucid, irregularly dentate, sometimes convolute, the apex muticous. Florets pale yellow. Achenes 4-5-5 mm, brown, with white ribs; pappus 5-7 mm. River sands. • S.E. Ukraine (by west bank of R. Dnepr, below Zaporož'e).