Centaurea margaritacea subsp. margaritacea

Tutin, T. G., Heywood, V. H., Burges, N. A., Moore, D. M., Valentine, D. H., Walters, S. M. & Webb, D. A., 1976, Flora Europaea. Volume 4. Plantaginaceae to Compositae (and Rubiaceae), Cambridge University Press : 286

(c) Subsp. margaritacea :

Stems scabrid above. Leaf-segments linear. Involucre 15-18 x 20-22 mm, depressed-globose; appendages 6-7 x 7-9 mm, not completely covering the yellow bracts, orbicular, convex, keeled, not pellucid, irregularly denticulate, convolute, the apex muticous. Florets yellowish or white. Achenes 3-5-4-5 mm, pale brown; pappus as long as or somewhat longer than achene. Submaritime sands. • S. Ukraine (W. side of estuary of R. Bug).