Centaurea margaritacea subsp. pineticola

Tutin, T. G., Heywood, V. H., Burges, N. A., Moore, D. M., Valentine, D. H., Walters, S. M. & Webb, D. A., 1976, Flora Europaea. Volume 4. Plantaginaceae to Compositae (and Rubiaceae), Cambridge University Press : 286

(j) Subsp. pineticola (Iljin) Dostál, loc. cit. (1976)

( C. pineticola Iljin ):

Stems scabrid above. Leaf-segments narrowly oblong. Involucre 15-18 x 10-14 mm, ovoid; appendages 4-6 mm, not covering bracts, orbicular, white, soft, indistinctly keeled, with triangular, dark blackish-brown central spot, the margin subdenticulate, pellucid, flat, the apex muticous. Florets pale purple. Achenes c. 4 mm, blackish; pappus as long as or somewhat longer than achene. Sandy pine woods. • S.C. Russia (S.E. of Voronež).