Centaurea margaritacea subsp. protogerberi

Tutin, T. G., Heywood, V. H., Burges, N. A., Moore, D. M., Valentine, D. H., Walters, S. M. & Webb, D. A., 1976, Flora Europaea. Volume 4. Plantaginaceae to Compositae (and Rubiaceae), Cambridge University Press : 286

(h) Subsp. protogerberi (Klokov) Dostál, loc. cit. (1976)

( C. protogerberi Klokov ):

Stems scabrid above. Involucre 14-15 x 12—14 mm, ovoid-globose; appendages 5 x 5-6 mm, not covering bracts, orbicular-ovate, yellow, hard, convex, indistinctly keeled, with triangular, subcoriaceous centre, the margin narrow, pellucid, flat, the apex muticous. Florets pale pink, yellowish or white. Achenes 4-5 mm, blackish-brown; pappus as long as achene. River sands. • Borders of S. Russia and E. Ukraine (by R. Donets).