Centaurea margaritacea subsp. donetzica

Tutin, T. G., Heywood, V. H., Burges, N. A., Moore, D. M., Valentine, D. H., Walters, S. M. & Webb, D. A., 1976, Flora Europaea. Volume 4. Plantaginaceae to Compositae (and Rubiaceae), Cambridge University Press : 286

(i) Subsp. donetzica (Klokov) Dostál, loc. cit. (1976)

( C. donetzica Klokov ):

Stems scabrid above. Segments of basal leaves broadly linear, of upper leaves narrowly linear. Involucre 12-15 x 10-16 mm, globose; appendages 5-6 x 4-4-5 mm, not covering bracts, elliptical, soft, indistinctly keeled, with shortly triangular, reddish-brown basal spot, the margin white, pellucid, shining, flat, the apex muticous. Florets pink. Achenes 4-5 mm, pale brown; pappus as long as achene. River sands. • E. Ukraine and S. Russia (Donets valley).