Amblygnathus puncticollis (Putzeys)

Shpeley, Danny, Hunting, Wesley & Ball, George E., 2017, A taxonomic review of the Selenophori group (Coleoptera, Carabidae, Harpalini) in the West Indies, with descriptions of new species and notes about classification and biogeography, ZooKeys 690, pp. 1-195 : 19

Amblygnathus puncticollis (Putzeys) Figs 5B, 6 D–F, 7B, 8

Selenophorus puncticollis Putzeys, 1878a: 34. LECTOTYPE male, labelled: St. Domingo [green paper, handwritten]; Soc. Ent. Belg. Coll. Putzeys; det. Putzeys Selenophorus puncticollis Put.; Type [red print]; Amblygnathus puncticollis Putz. V. Emd. Det. 1937; R.I.Sc.N.B.I.G. (IRSB).- Csiki 1932: 1200; Darlington 1934: 104.- Blackwelder 1944: 50.

Amblygnathus puncticollis ; Erwin and Sims 1984: 441.- Ball and Maddison 1987: 223.- Ball 1992: 85.- Lorenz 1998: 356.- Lorenz 2005: 378.- Perez-Gelabert 2008: 79.

Type area.

"Santo Domingo" = Greater Antillean island of Hispaniola.

Diagnosis.

This species is readily separated from the other two West Indian Amblygnathus species on the basis of small size and its range restricted to the Greater Antilles.

Descriptive notes.

Data for SBL in Table 1. Habitus as in Fig. 5B. Males with two terminal setae and females with four terminal setae near the posterior margin on sternum VII.

Male genitalia. Figs 6 D–F. Apical portion of phallic median lobe moderate in length, trapezoidal and broadly rounded in dorsal aspect, with very narrow dorsal flange visible only laterally on both sides; endophallus with two moderately long spines and two darkened microtrichial fields; lamina present, short, broad at base, tapered sharply at apex. A single male from San Vicente in Cuba has the apical portion of the phallic median lobe with a fully developed dorsal flange and the spines and michrotrichial fields of the endophallus are a bit differently oriented. At this time, we prefer to consider this a variant rather than a different species.

Ovipositor and female reproductive tract. Fig. 7B. Gonocoxite 2 falcate with moderately wide base. Bursa copulatrix moderately long; spermatheca (sp) long, loosely coiled, broadly attached near the base of the common oviduct. Spermathecal gland duct originating above the base of the spermatheca, spermathecal gland (spg) small, sausage-like, long double swelling of duct basad gland.

Geographical distribution.

Fig. 8. This species is known only from the Greater Antillean islands of Cuba, Hispaniola, Jamaica and Puerto Rico.

Chorological affinities and relationships.

The range of this species is not overlapped by the other West Indian species of Amblygnathus . Its putative adelphotaxon is the Middle American A. woodruffi Ball & Maddison. (See Ball and Maddison 1987: 261, Fig. 70B).

Material examined.

In addition to type material, we have seen a total of 19 specimens (12 males, 7 females). See Appendix for details.