Dryininae Haliday, 1833
publication ID |
https://doi.org/ 10.11646/zootaxa.3614.1.1 |
publication LSID |
lsid:zoobank.org:pub:E75E5224-20F1-431C-A7CB-9EE0D3F25118 |
DOI |
https://doi.org/10.5281/zenodo.6498554 |
persistent identifier |
https://treatment.plazi.org/id/8860B757-8EA1-FF31-48A7-5024FE38FE22 |
treatment provided by |
Felipe |
scientific name |
Dryininae Haliday, 1833 |
status |
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V. Subfamily Dryininae Haliday, 1833
Dryini partim Haliday 1833: 273.
Dryininae partim Kieffer in Kieffer & Marshall 1906: 495.
Dryininae partim Kieffer 1907: 3.
Lestodryinini partim Kieffer 1914b: 11.
Dryinini partim Richards 1939: 189. Dryinini partim Muesebeck & Walkley 1951: 1034.
Dryininae Richards 1953: 51 ; Olmi 1984: 723; Olmi 1993a: 194; Olmi 1993b: 59; Olmi 1993d: 46; Olmi 1994: 62; Olmi 1995b: 503; Olmi 1999a: 180; He & Xu 2002: 242; Engel 2005: 486; Olmi & Virla 2006: 408; Virla & Olmi 2008: 370.
Dryininae partim Krombein 1979: 1243.
Thaumatodryininae partim Olmi 1984: 682.
Thaumatodryininae partim Olmi 1993b: 59.
Type genus. Dryinus Latreille, 1804 .
Diagnosis. Female ( Plates 73A View PLATE 73 , 83A View PLATE 83 ): fully winged ( Plates 73A View PLATE 73 , 83A View PLATE 83 ); ocelli present; occipital carina complete, or incomplete, or absent; palpal formula usually 6/3; occasionally palpal formula different (4/ 2 in Gonadryinus Olmi ; 3/2, or 4/2, or 5/ 3 in Pseudodryinus Olmi ); mandible usually with four teeth progressing larger from anterior one to posterior (as in plate 3D); occasionally (in Pseudodryinus Olmi ) mandible with four irregular teeth; antenna usually without tufts of long hairs ( Plate 83A View PLATE 83 ); occasionally (in Thaumatodryinus R. Perkins ) antennal segments 5–10 with tufts of long hairs ( Plates 69E View PLATE 69 , 70A View PLATE 70 , 73A View PLATE 73 ); pronotal tubercle present ( Plate 73A View PLATE 73 ); epicnemium visible, because lateral regions of prothorax not continuous with mesopleura (as in plate 2H); fore wing with three cells enclosed by pigmented veins (costal, median and submedian) ( Plates 73A View PLATE 73 , 83A View PLATE 83 ); occasionally (in Thaumatodryinus R. Perkins ) also first brachial cell clearly enclosed by pigmented veins ( Plate 69E View PLATE 69 ); fore wing with stigmal vein and pterostigma ( Plates 73A View PLATE 73 , 83A View PLATE 83 ); protarsus chelate ( Plates 69E View PLATE 69 , 73A View PLATE 73 , 83A View PLATE 83 ); chela with rudimentary claw ( Plates 73B View PLATE 73 , 83A View PLATE 83 ); protrochanter long and slender ( Plate 83A View PLATE 83 ); usually tibial spurs 1/1/2; rarely (in some species of Dryinus Latreille ) 1/1/1. Male ( Plates 72A View PLATE 72 , 76A View PLATE 76 ): fully winged ( Plates 72A View PLATE 72 , 76A View PLATE 76 ); palpal formula 6/3; occipital carina complete or incomplete; mandible usually with one–three teeth (three teeth progressing larger from anterior one to posterior); occasionally (in Thaumatodryinus R. Perkins ) mandible with four teeth progressing larger from anterior one to posterior (as in plate 3D); occasionally (in Pseudodryinus Olmi ) mandible with four irregular teeth; epicnemium visible, because lateral regions of prothorax not continuous with mesopleura ( Plate 2H View PLATE 2 ); mesosternum fused with mesopleura and not distinct; fore wing with three cells enclosed by pigmented veins (costal, median and submedian) ( Plates 72A View PLATE 72 , 76A View PLATE 76 ); fore wing with stigmal vein and pterostigma ( Plates 72A View PLATE 72 , 76A View PLATE 76 ); fore wing with metacarpus as long as, or longer than pterostigma ( Plates 72A View PLATE 72 , 76A View PLATE 76 ); dorsal process of paramere absent ( Plates 71A View PLATE 71 , 74C View PLATE 74 ); tibial spurs 1/1/2.
Distribution. Worldwide.
Hosts. Acanaloniidae , Cixiidae , Dictyopharidae , Flatidae , Fulgoridae , Issidae , Lophopidae , Ricaniidae , Tropiduchidae ( Guglielmino & Olmi, 1997, 2006, 2007).
World species. Three hundred and fourty two species are known, sixty three in the Oriental region.
World genera. Nine genera known, four in the Oriental region.
Remarks. Haliday is considered author of Dryininae for the principle of coordination (Art. 36 ICZN). In fact, the author of the family is also author of the subfamily whose name derives from the family name. This happens even if Kieffer was the first who introduced the subfamily division.
The following five genera are not quoted from the Oriental region: Cretodryinus Ponomarenko, 1975d (fossil in Taimyr amber), Gonadryinus Olmi, 1991 , Harpactosphecion Haupt, 1944 (fossil in amber from the Dominican Republic and Baltic area), Megadryinus Richards, 1953 (Neotropical) , Palaeodryinus Olmi & Bechly, 2001 (fossil in Baltic amber).
Key to the genera (excluding Hybristodryinus Engel )
Females
1. Palpal formula 3/2, 4/2, 5/2, 4/3, or 5/3.................................................. 3. Pseudodryinus Olmi
- Palpal formula 6/3.....................................................................................2
2. Antenna with tufts of long hairs on segments 5–10 ( Plates 69E View PLATE 69 , 70A View PLATE 70 , 73A View PLATE 73 ).................. 1. Thaumatodryinus Perkins
- Antenna without tufts of long hairs ( Plate 83A View PLATE 83 )............................................... 2. Dryinus Latreille
Males
1. Mandible with one–three teeth............................................................ 2. Dryinus Latreille
- Mandible with four teeth.................................................................................2
2. Notauli always separated at posterior margin of scutum ( Plate 72A View PLATE 72 ); minimum distance between notauli as long as, or longer than greatest breadth of posterior ocelli ( Plate 72A View PLATE 72 )...................................1. Thaumatodryinus R. Perkins
- Notauli joint or separated at posterior margin of scutum; if separated, minimum distance between notauli much shorter than greatest breadth of posterior ocelli.......................................................3. Pseudodryinus Olmi
Remarks. Hybristodryinus Engel, 2005 , is based on one only species, resinicolus Engel, 2005 . Its holotype (only known specimen) is in very bad conditions, so that it is impossible to find characters useful for including it in the above key to the genera. For this reason it is left separated from the other three genera and put at the end of the treatment of Dryininae .
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
Kingdom |
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Order |
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SuperFamily |
Chrysidoidea |
Family |
Dryininae Haliday, 1833
Xu, Zaifu, Olmi, Massimo & He, Junhua 2013 |
Thaumatodryininae
Olmi, M. 1993: 59 |
Thaumatodryininae
Olmi, M. 1984: 682 |
Dryininae
Krombein, K. V. 1979: 1243 |
Dryininae
Virla, E. G. & Olmi, M. 2008: 370 |
Engel, M. S. 2005: 486 |
He, J. & Xu, Z. 2002: 242 |
Olmi, M. 1999: 180 |
Olmi, M. 1994: 62 |
Olmi, M. 1993: 194 |
Olmi, M. 1993: 59 |
Olmi, M. 1993: 46 |
Olmi, M. 1984: 723 |
Richards, O. W. 1953: 51 |
Olmi 1995b: 503 |
Olmi & Virla 2006: 408 |
Dryinini
Muesebeck, C. F. W. & Walkley, L. M. 1951: 1034 |
Richards, O. W. 1939: 189 |
Dryininae
Kieffer, J. J. 1907: 3 |
Dryininae
Kieffer, J. J. & Marshall, T. A. 1906: 495 |
Dryini partim Haliday 1833: 273 |
Lestodryinini partim Kieffer 1914b: 11 |