Dobarakuku, Perlaza-Cruz & Espinosa & Albo & Labarque & Cabra-García, 2025
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publication ID |
https://doi.org/10.11646/zootaxa.5725.3.6 |
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publication LSID |
lsid:zoobank.org:pub:F280069F-3587-4D87-97A5-DFFE633A9669 |
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DOI |
https://doi.org/10.5281/zenodo.17869150 |
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persistent identifier |
https://treatment.plazi.org/id/830987AE-D92D-FF9A-3ACB-FEB3FB7BE343 |
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treatment provided by |
Plazi |
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scientific name |
Dobarakuku |
| status |
gen. nov. |
Dobarakuku gen. nov.
Type species. Tinus ursus Carico, 1976 View in CoL . Designated here.
Etimology. The generic name is a noun taken from the Emberá languages that means “river spider” ( do = river; barakuku = spider). Gender is masculine.
Diagnosis. Males can be distinguished from all other Thaumasiinae genera by the following combination of characters: narrow DTP occupying less than half the width of the cymbial alveolus ( Fig. 3A View FIGURE 3 ), longer in Architis ( Santos 2007b: fig. 2A) and Thaumasia ( Silva & Carico 2012: fig. 34), and absent in Tinus ( Carico 1976: fig. 8); tegulum with PTP ( Fig. 4A, C View FIGURE 4 ), absent in Architis ( Sierwald 1990: fig. 23); DST with its basal part exposed at the prolateral margin of the tegulum in ventral view ( Fig. 3A View FIGURE 3 ), absent in Architis ( Sierwald 1990: fig. 23) and Thaumasia ( Sierwald 1990: fig. 18); laminar, hyaline conductor covering the embolus tip ( Fig. 3A View FIGURE 3 ), spatulated in Tinus ( Carico 1976: fig. 12) and Thaumasia ( Silva & Carico 2012: fig. 145); vestigial MA ( Fig. 3A View FIGURE 3 ), well-developed in Thaumasia ( Silva & Carico 2012: fig. 141); and stout embolus ( Fig. 4A, C View FIGURE 4 ), filiform in Architis ( Santos 2007b: fig. 4A), Thaumasia ( Silva & Carico 2012: fig. 140) and Tinus ( Carico 1976: fig. 16), basally fused with the DST ( Fig. 4A, C View FIGURE 4 ). Females can be diagnosed by the following combination of characters: prominent lateral lobes of the epigynum projecting posteriorly ( Fig. 3C, D View FIGURE 3 ), smaller in Architis ( Santos 2007b: fig. 16C) and Thaumasia ( Silva & Carico 2012: fig. 68); epigynum with two conspicuous oval atria ( Fig. 3C, D View FIGURE 3 ), when present, smaller in Architis ( Santos 2007b: fig. 3C); and elongated spermathecae head ( Fig. 3D View FIGURE 3 ), subspherical in Tinus ( Carico 1976: fig. 23).
Description. Medium sized nursery-web spiders, with males (total length 6.95–9.20) of similar size as females (total length 7.60–10.45). Carapace longer than wide, oval, narrowest anteriorly and widest between coxae II and III ( Fig. 2A, D View FIGURE 2 ). Colouration yellowish-brown, with two lateral longitudinal pale stripes with irregular borders, extending from the anterior to the posterior edge, and a narrow light stripe along the midline ( Fig. 2A, D View FIGURE 2 ). Covered with dark setae ( Fig. 2C, F View FIGURE 2 ). Ocular area darker, eyes encircled by black rings ( Fig. 2A, D View FIGURE 2 ). Thoracic fovea longitudinal and distinct. Anterior eye row straight in dorsal view; posterior eye row recurved. Median eyes larger than laterals. Clypeus height approximately 0.7 to 1.3 times anterior median eyes diameter. Chelicerae with two to three promarginal and three retromarginal teeth. Endites rectangular, with anteromedian scopula apically ( Fig. 2B, E View FIGURE 2 ). Labium wider than long, distally truncated, and laterally notched in the proximal half ( Fig. 2B, E View FIGURE 2 ). Sternum wider than long, posteriorly triangular, with posterior tip projecting between coxae IV ( Fig. 2B, E View FIGURE 2 ). Leg tarsi with apical pseudosegment. Opisthosoma pilose, oval to cylindrical, longer than wide ( Fig. 2A, D View FIGURE 2 ). Colouration dark yellow to dark brown marked with irregular light spots, plus two yellowish longitudinal stripes along the lateral margins, resembling those on the carapace, and two additional longitudinal stripes positioned closer to the medial region, converging near the posterior edge ( Fig. 2A, D View FIGURE 2 ). Venter usually paler than dorsum ( Fig. 2B, E View FIGURE 2 ). Six spinnerets, posterior laterals longer and narrower than anterior laterals.
Palp with retrolateral tibial apophysis conspicuous and bifurcated ( Fig. 3A View FIGURE 3 ). The dorsal branch is thick, broad, and highly sclerotized at the tip, while the ventral branch is slender, curved, and less sclerotized at the tip ( Fig. 3A View FIGURE 3 ). The cymbium presents a well-developed basal apophysis with rounded edges ( Fig. 3A View FIGURE 3 ). The basal hematodocha expands prominently during the palp expansion ( Fig. 4 View FIGURE 4 ). The tegulum consists of two distinct portions. The basal portion is ring-shaped and strongly sclerotized, bearing a proximal tegular projection (PTP) near the point where the sperm duct enters the tegulum ( Fig. 4A, C View FIGURE 4 ). The PTP is not visible in the unexpanded palp as it is covered by the distal tegular projection (DTP) ( Fig. 3A View FIGURE 3 ). The distal portion of the tegulum is strongly sclerotized at its posterior border and membranous in its medial area ( Fig. 3A View FIGURE 3 ). This portion bears the conductor, the median apophysis, and the DTP ( Fig. 3A View FIGURE 3 ). The conductor is hyaline and extends apically, covering the embolus tip in the unexpanded palp ( Fig. 3A View FIGURE 3 ). The median apophysis is vestigial ( Fig. 3A View FIGURE 3 ). The DTP is highly sclerotized, prolaterally oriented, triangular, and has a rounded apical tip ( Fig. 3A View FIGURE 3 ). The sperm duct passes through the edges of the DTP, reaching its apical tip, where it makes a U-turn to transition into the apical division of the palp ( Fig. 4B View FIGURE 4 ). At this point, a basal membrane (BMT) connects the DTP to the distal sclerotized tube of the apical division (DST) ( Fig. 4A View FIGURE 4 ). The embolus is fused to the DST ( Fig. 4A, C View FIGURE 4 ). Its distal portion is flattened and curves retrolaterally in ventral view ( Figs 3A, B View FIGURE 3 , 4 View FIGURE 4 ).
Epigynum with a subtriangular posterior field and prominent lateral lobes projecting posteriorly ( Fig. 3C View FIGURE 3 ). Median field with a central elevation, separating two conspicuous oval atria ( Fig. 3C View FIGURE 3 ). Spermatheca positioned laterally, with slender heads, longer than their stalks. Copulatory ducts short, subequal in size to the fertilization ducts ( Fig. 3C, D View FIGURE 3 ).
Distribution. Known from the Pacific region of Costa Rica, Panama, and Colombia ( Fig. 5 View FIGURE 5 ).
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.