Boreophilia Benick, 1973

Klimaszewski, Jan, Sikes, Derek S., Brunke, Adam & Bourdon, Caroline, 2019, Species review of the genus Boreophilia Benick from North America (Coleoptera, Staphylinidae, Aleocharinae, Athetini): Systematics, habitat, and distribution, ZooKeys 848, pp. 57-102 : 57

Genus Boreophilia Benick, 1973 Figs 1, 2-5, 6-15, 16-23, 24-31, 32-42, 43-52, 53-56, 57-62, 63-70, 71-80, 81-83, 84-91, 92-99, 100-108, 109-116, 117-119

Boreophilia Benick, 1973: 211; Lohse et al. 1990: 151; Gusarov 2003a, b: 81; Smetana 2004: 396; Schülke and Smetana 2015: 557; Klimaszewski et al. 2018. Type species: Homalota islandica Kraatz, 1857.

Diagnosis.

Boreophilia may be distinguished from other athetines by the following combination of characters: body moderately narrow to broad, subparallel (Figs 6, 16, 24, 32, 43, 53, 57, 63, 71, 81, 84, 92, 100, 109, 117); head narrower or nearly as broad as pronotum (Fig. 6, 16, 24, 32, 43, 53, 57, 63, 71, 81, 84, 92, 100, 109, 117); eyes moderate in size, as long as or shorter than postocular region of head, tempora partially feebly carinate at base of head; antennae often reaching posterior margin of elytra, antennomeres V-X subquadrate, slightly transverse, or elongate (Figs 6, 16, 24, 32, 43, 53, 57, 63, 71, 81, 84, 92, 100, 109, 117); ligula divided into two separate and diverging lobes (Fig. 2); mandibles simple (not bifid at apex as in Schistoglossa ) (Figs 3, 4); maxillary palps with 4 palpomeres, terminal one needle-shaped (Fig. 5); pronotum transverse, broadest in the middle or in apical fourth, hind angles rounded, hypomeron clearly visible in lateral view; integument smooth and moderately glossy, weak microsculpture present, punctuation fine, pubescence on midline of disc directed anteriad at least in apical ¾, and elsewhere directed laterally; mesocoxae contiguous; median lobe of aedeagus broad, flattened latero-ventrally, bulbus enlarged and tubus narrow, short or moderately elongate, approximately triangularly shaped in dorsal view (Figs 9, 10, 18, 26, 35, 36, 46, 59, 60, 65, 74, 83, 86, 94, 102, 103, 111), crista apicalis narrowly elongate in most, internal sac with two large, elongate sclerites in bulbus, and additional smaller sclerites in apical part of internal sac; male tergite VIII entire, and not modified (Figs 11, 19, 27, 37, 47, 61, 66, 75, 87, 95, 104, 112); spermatheca diverse in shape, with elongate tubular capsule and sinuate stem, invagination of capsule small or absent (Figs 15, 23, 31, 41, 42, 51, 52, 56, 70, 79, 80, 91, 99, 108, 116, 120); cold loving species occurring in arctic habitats, in temperate regions usually confined to fens and bogs.