PUTOIDAE TANG, Tang

Moghaddam, Masumeh, 2013, A review of the mealybugs (Hemiptera: Coccoidea: Pseudococcidae, Putoidae and Rhizoecidae) of Iran, with descriptions of four new species and three new records for the Iranian fauna, Zootaxa 3632 (1), pp. 1-107 : 94

FAMILY PUTOIDAE TANG

Recent phylogenetic analyses based on nucleotide sequence data have shown that the genus Puto does not belong to the Pseudococcidae (Cook et al. 2002; Gullan & Cook, 2007; Hardy et al. 2008) but should be placed in a separate family, Putoidae , as indicated by Williams et al. (2011, 2008).

DIAGNOSIS. Adult female: eyes tall, height usually as great as length of first antennal segment; antennae usually 9 segmented, rarely 8; antennal intersegmental sensilla present between segments III & IV, IV & V and VI & VII, never present between segments V & VI; each surface of trochanter with 2–5 (usually 3 or 4) campaniform sensilla; claw usually with a pair of basal spurs; claw digitules often capitate but tarsal digitules almost never capitate; with at least 18 pairs of cerarii on sclerotised plates, if more pairs present, increase due to division of cerarii on certain segments; long tubular ducts almost always present on frons (anterior to mouthparts); multilocular pores present on venter; quinquelocular pores absent; trilocular pores usually of 3 sizes, ventral pores smallest, dorsal pores noticeably larger, cerarian pores slightly larger than dorsal pores.