Bipectinata orientalis, Wichard & Espeland & Müller & Wang, 2020

Wichard, Wilfried, Espeland, Marianne, Müller, Patrick & Wang, Bo, 2020, New species of caddisflies with bipectinate antennae from Cretaceous Burmese amber (Insecta, Trichoptera: Odontoceratidae, Calamoceratidae), European Journal of Taxonomy 653, pp. 1-17 : 13-15

Bipectinata orientalis gen. et sp. nov.

urn:lsid:zoobank.org:act:988D3108-67F2-473D-85AB-E21A4E02DA46

Fig. 5

Diagnosis

See the corresponding diagnosis of the genus

Etymology

The species name describes the biogeographic region in which this new fossil species was found.

Material examined

Holotype

MYANMAR – Kachin State • ♂; exact locality unknown; Mid-Cretaceous Burmese amber inclusion; NIGP172206 .

Description

HEAD. Antennae about as long as forewings, partially bipectinate, first consisting of long scapus, short pedicellus and simple flagellomere longer than pedicellus, both together about as long as scapus; followed by 16 successive bipectinate flagellomeres and, finally, 18 simple flagellomeres without rami. Rami originating on both sides at base of flagellomere, basally slender and gradually expanding towards rounded apex. Maxillary palps 5-segmented, labial palps 3-segmented, their terminal segments not flexible or annulate. Head dorsally with broad posterior setose warts, transverse, covered with single, long hairs; scutum densely haired, setose warts not visible, scutellum shows single dome-shaped, large, ovoid wart.

FOREWINGS. Light brown, slightly translucent, 6 mm long. Radius R1 running straight to apical wing margin. Forks I and II of Rs present, sessile, branching at same level. Nygma at base of fork II. Discoidal cell closed by crossveins, about as long as its stem Rs. Media branching first into M1+2 and M3+4 and then forks III and IV, medial cell absent or open. Cu1 running straight, apically bifurcated into fork V (Cu1a–Cu1b); crossvein m-cu between Cu1a and M4, therefore thyridial cell present, very probably with nygma. Cu2 simple, not forked. Anal veins probably complete. Hindwings ( Fig. 5C ) light brown, translucent, smaller than forewings, about 4.5 mm long, Radius R1 straight, forks I, II present, fork II with nygma basally; discoidal cell absent or open. M branched in fork III and in M3+4. Cu1 with fork V, Cu2 simple. Crossveins not clearly visible, but indicated in forewings through fine, small hairs, running parallel to each other and forming ‘staggered bridge’ between longitudinal veins.

TIBIAL SPURS. 2/4/4.

MALE GENITALIA. Visible only in ventral view: Inferior appendages (gonocoxites) 2-segmented consisting of straight digitiform coxopodite and curved rod-shaped harpago, length ratio 3:1; harpago initially thin and gradually enlarging into small club. Pair of long needles situated between inferior appendages, slightly dorsally placed and curved medio-dorsad, sitting on slightly stronger pedestal. In ventral view central phallic apparatus is possibly covered by 2 membranous dorsal scales, probably derived from the Xth segment, and laterally flanked by 2 short parameres [phallic apparatus itself remains unclear].

Figures

Fig. 5. Bipectinata orientalis sp. nov. (NIGP172206) A. Holotype male in ventral view. B. Bipectinate antennae with scapus, pedicellus and flagellomeres. C. Drawing of fore- and hindwings. D. Drawing of the male genitalia. E. Male genitalia in dorsal view.