Megalyrinae Schletterer, 1889, 2024

Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo & Perrichot, Vincent, 2024, Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities, Arthropod Systematics & amp; Phylogeny 82, pp. 151-181 : 151

3.2. Subfamily Megalyrinae Schletterer, 1889

Emended diagnosis.

Head globular; compound eye oval, with or without postocular carina; flagellomeres often elongate, sometimes compact; subantennal groove with or without dorsal carina, mandibles symmetrical with three teeth. Anterior thoracic spiracle fully surrounded by pronotal cuticle; median mesoscutal sulcus sometimes effaced or reduced; parapsidal lines sometimes absent. Fore wing with infumate banding patterns in Ettchellsia , Dinapsis , Megalyra and Megalyridia ; at least Sc+R, Rs+M, basal segment of Rs and basal segment of R1 pigmented and no closed cell; at most with C, Sc+R, A, cu-a, Rs+M, Rs, R1 and basal segment of M pigmented, costal, submarginal, and marginal cells closed by tubular veins, subcostal and first cubital fused; M+Cu usually absent except in † Cretodinapsis ; rarely brachypterous. Hind leg without row of erect setae along margin. Metasoma usually elongate but sometimes distinctly shorter than mesosoma.

Genera included.

Carminator Shaw, 1988, Cryptalyra Shaw, 1987 [ Cryptalyrini Shaw, 1988]; Dinapsis Waterston, 1922, Ettchellsia Cameron, 1909, Neodinapsis Shaw, 1987 [ Dinapsini Waterston, 1922]; † Cretodinapsis Rasnitsyn, 1977, Megalyra Westwood, 1832, † Prodinapsis Brues, 1923 [ Megalyrini Schletterer, 1889]; † Kamyristi gen. nov. [† Kamyristini trib. nov]; Megallica Perrichot, 2009 [† Megallicini trib. nov.]; Megalyridia Hedqvist, 1959 [ Megalyridiini Shaw, 1990]; Rigel Shaw, 1987 [ Rigelini Shaw, 1990]; † Valaa Perrichot, 2009 [† Valaaini trib. nov.].

Stratigraphic extension.

Albian to present.

Comments.

We redefine the subfamily Megalyrinae , according to the results of the phylogenetic analyses. The subfamily is composed of three non-monotypic tribes, namely Cryptalyrini , Dinapsini , and Megalyrini , as well as a number of monotypic tribes: Rigelini , † Kamyristini trib. nov., † Megallicini trib. nov., † Valaaini trib. nov., and Megalyridiini . The justification for redefining Megalyrinae is given in the Discussion.