Ummidia rodeo, Godwin & Bond, 2021

Godwin, Rebecca L. & Bond, Jason E., 2021, Taxonomic revision of the New World members of the trapdoor spider genus Ummidia Thorell (Araneae, Mygalomorphae, Halonoproctidae), ZooKeys 1027, pp. 1-165 : 1

Ummidia rodeo sp. nov. Figs 37 , 38 , Map 4

Type material.

HOLOTYPE ♂ (UMM325) from 5 miles south of Rodeo, Durango, Mexico, 25.1081 -104.5556 5, 1365 m a.s.l., 17.vii.1975, coll. D Ubick, AMNH. PARATYPE: 1 ♀ (UMM166) from Yerbanís, 80 miles northeast of Durango, 24.7356 -103.8364 6, 1905 m a.s.l., 1.viii.1947, coll. WJ Gertsch, AMNH.

Etymology.

The specific epithet is a noun taken in apposition and is in reference to the type locality.

Diagnosis.

Ummidia rodeo males can be differentiated from U. gabrieli by the presence of prolateral spines and by having relatively more retrolateral spines (24 vs 11) on tibia I. Females can be differentiated from U. gabrieli by having relatively more distal endite cuspules (59 vs 28), from U. pustulosa by have the sternum and carapace roughly as wide as long, and from all geographically proximate species by having the labium 2 × wide as long (1.5 × wide as long in other species). Males disperse in July.

Description of male holotype.

Specimen preparation and condition. Specimen preserved in 80% EtOH. Left palp and legs I&II removed; in vial with specimen. General coloration. Carapace and chelicerae reddish black 7.5YR 2.5/1, legs very dusky red 2.5YR 2.5/2. Abdomen black 10YR 2/1. Cephalothorax. Carapace 7.20 long, 6.95 wide. Pars cephalica 5.05 long. Foveal groove procurved, 0.46 long, 1.55 wide. Tubercle moderate, only under AME. AER slightly procurved. PER relatively straight. Eye group 0.74 long, 1.46 wide, AME 0.38, PME 0.17, ALE 0.36, PLE 0.19. Sternum sparsely setose, STRl 3.92, STRw 3.84. Chelicerae with anterior tooth row comprising six teeth, posterior margin with four teeth. Palpal endites with 13 non-hastate cuspules over proximal 1/4 of endite face, lacking distal endite cuspules, ENDw 1.64, ENDl 2.88. Labium with seven small cuspules, LBw 1.48, LBl 1.23. Rastellum with five small spines along distal margin of process. Abdomen setose. Legs. F1 6.42; F1w 1.88; P1 3.01; Ti1 4.28; Mt1 2.97; Tr1 1.53; F3 4.84; F3w 2.16; P3 2.55; Ti3 2.86; Sd3 2.11; Mt3 2.7; Tr3 1.74; F4 6.45; F4w 2.03; P4 2.92; Ti4 3.99; Mt4 4.33; Tr4 1.85. Retrolateral face of tarsus IV lacking distinct comb or brush. Leg I spination pattern: TSp 16, TSpv 10, TSrd 0, TSr 0, TSrv 24, MtSp 2, MtSr 9, TrSp 1, TrSr 13. Pedipalps. PTl 3.83, PTw 1.36, Bl 3.43. Embolus sinuous.

Variation, males

(n = 2). CL 6.17-7.2, 6.68 ± 0.52; CW 6.22-6.95, 6.58 ± 0.36; Cap 4.19-5.05, 4.62 ± 0.43; ENDl 0.65-0.74, 0.69 ± 0.04; ENDw 1.46-1.48, 1.47 ± 0.01; STRl 3.59-3.92, 3.75 ± 0.17; STRw 3.33-3.84, 3.58 ± 0.26; LBl 0.77-1.64, 1.21 ± 0.44; LBw 1.06-1.48, 1.27 ± 0.21; F1 5.79-6.42, 6.1 ± 0.32; F1w 1.46-1.88, 1.67 ± 0.21; P1 2.64-3.01, 2.83 ± 0.18; Ti1 3.48-4.28, 3.88 ± 0.4; Mt1 2.52-2.97, 2.75 ± 0.23; Tr1 1.52-1.53, 1.53 ± 0; F3 4.13-4.84, 4.48 ± 0.35; F3w 1.68-2.16, 1.92 ± 0.24; P3 2.24-2.55, 2.39 ± 0.16; Ti3 2.11-2.24, 2.17 ± 0.06; Mt3 2.49-2.7, 2.6 ± 0.1; Tr3 1.74-1.83, 1.78 ± 0.04; F4 5.23-6.45, 5.84 ± 0.61; F4w 1.73-2.03, 1.88 ± 0.15; P4 2.47-2.92, 2.7 ± 0.23; Ti4 3.35-3.99, 3.67 ± 0.32; Mt4 3.5-4.33, 3.92 ± 0.41; Tr4 0-1.94, 0.97 ± 0.97; TSp 10-16, 13 ± 3; TSpv 5-10, 7.5 ± 2.5; TSr 0-0, 0 ± 0; TSrv 0-23, 11.5 ± 11.5; PTl 3.74-3.83, 3.78 ± 0.05; PTw 1.23-1.36, 1.29 ± 0.06; BL 2.95-3.43, 3.19 ± 0.24.

Description of female paratype.

Specimen preparation and condition. Specimen preserved in 80% EtOH. Spermathecae removed, cleared, in vial with specimen. General coloration. Carapace, chelicerae, and legs very dusky red 10R 2.5/2. Abdomen reddish black 2.5YR 2.5/1, spinnerets dark yellowish brown 10YR 3/6. Cephalothorax. Carapace 9.26 long, 8.57 wide. Pars cephalica 6.39 long. Foveal groove procurved, 0.67 long, 2.39 wide. Eye tubercle low. AER procurved. PER straight to slightly recurved. Eye group 1.02 long, 2.08 wide, AME 0.43, PME 0.22, ALE 0.52, PLE 0.33. Sternum sparsely setose, STRl 5.95, STRw 5.24. Chelicerae with anterior row comprising seven teeth, posterior margin with six teeth. Palpal endites with 15 cuspules spread across proximal half of endite and 42 smaller cuspules distally, ENDw 2.08, ENDl 3.45. Labium with three cuspules, LBw 2.07, LBl 1.08. Rastellum with many strong spines on process and up cheliceral face for ~ 1 × length of process. Abdomen. Evenly setose with pale speckles clustered at apodemes. Legs. F1 5.1; F1w 1.9; P1 3.2; Ti1 3.04; Mt1 2.3; Tr1 1.25; F3 4.7; F3w 2.6; P3 2.85; Ti3 2.56, Sd3 1.55; Mt3 2.03; Tr3 1.86; F4 6.12; F4w 2.65; P4 3.08; Ti4 3.12; Mt4 3.33; Tr4 1.82. Retrolateral face tarsus IV with defined comb over length of tarsus. Pedipalps. PF 4.75, PP 2.54, PTi 3.1, PTr 2.6. Spermathecae curved laterally with distal coil, bulbs face anterolaterally.

Variation, females

(n = 5). CL 8.23-9.26, 8.72 ± 0.17; CW 7.74-8.57, 8.15 ± 0.14; Cap 5.77-6.39, 6.02 ± 0.11; ENDl 0.86-1.04, 0.98 ± 0.03; ENDw 1.17-1.87, 1.65 ± 0.13; STRl 5.05-5.95, 5.37 ± 0.16; STRw 4.58-5.24, 5.01 ± 0.13; LBl 1.08-1.37, 1.22 ± 0.05; LBw 1.81-2.07, 1.96 ± 0.04; F1 4.69-5.1, 4.85 ± 0.07; F1w 1.73-1.9, 1.83 ± 0.03; P1 2.97-3.2, 3.1 ± 0.04; Ti1 2.88-3.06, 2.96 ± 0.04; Mt1 1.96-2.3, 2.08 ± 0.07; Tr1 1.15-1.44, 1.27 ± 0.05; F3 4.28-4.75, 4.55 ± 0.09; F3w 2.35-2.62, 2.48 ± 0.05; P3 2.85-2.95, 2.89 ± 0.02; Ti3 2.48-2.7, 2.59 ± 0.04; Mt3 1.77-2.03, 1.88 ± 0.05; Tr3 1.72-1.95, 1.84 ± 0.05; F4 5.59-6.12, 5.78 ± 0.1; F4w 2.41-2.65, 2.52 ± 0.05; P4 2.81-3.31, 3.04 ± 0.08; Ti4 2.92-3.26, 3.08 ± 0.06; Mt4 2.59-3.33, 3.06 ± 0.13; Tr4 1.63-1.91, 1.75 ± 0.05; PF 4.54-4.75, 4.63 ± 0.04; PP 2.54-2.78, 2.63 ± 0.04; PTi 2.93-3.19, 3.06 ± 0.05; PTr 2.44-2.69, 2.57 ± 0.04.

Material examined.

Mexico: Chihuahua: 5 mi E Parral , 26.9323 -105.5847 5, 548 m a.s.l. (UMM0626, 15.vii.1947, 1♀, WJ Gertsch, AMNH); Summit NE of San José Babícora, 29.2979 -107.537 5, 1158 m a.s.l. (UMM0629, 3.vii.1947, 1♂, WJ Gertsch, AMNH) ; Durango: San Lucas Ocampo , 24.0072 -104.6662 6, 1887 m a.s.l. (UMM0298, 1.viii.1947, 1♀, WJ Gertsch, AMNH); Yerbanís, 80 mi NE Durango, 24.7356 -103.8364 6, 1905 m a.s.l. (UMM0444, 19.viii.1947, 1♀, WJ Gertsch, AMNH); (UMM0308, 1♀, AMNH); (UMM0448, 1♀, AMNH); (UMM0449, 1♀, AMNH); (UMM0581, 1♀, AMNH); (UMM0619, 1♀, AMNH); (UMM0166, 1♀, AMNH); 5 mi S Rodeo, 25.1081 -104.5556 5, 1365 m a.s.l. (UMM0325, 17.vii.1975, 1♂, D Ubick, AMNH) .

Figures

Map 4. Locality records for western United States and northern Mexico species.

Figure 37. Ummidia rodeo sp. nov. from Durango, Mexico A male habitus illustration UMM 325 B female habitus illustration UMM 166. Scale bars: 4.0 mm.

Figure 38. Ummidia rodeo sp. nov. from Durango, Mexico A-D male holotype (UMM 325) A prolateral aspect, leg I B retrolateral aspect, leg I C line drawings, leg I prolateral and retrolateral aspects D retrolateral aspect, pedipalp E cleared spermathecae female paratype (UMM 166). Scale bars: 1.0 mm.