Exocelina miriae (Balke, 1998)

Shaverdo, Helena, Sagata, Katayo & Balke, Michael, 2016, Taxonomic revision of New Guinea diving beetles of the Exocelinadanae group, with the description of ten new species (Coleoptera, Dytiscidae, Copelatinae), ZooKeys 619, pp. 45-102 : 48

Taxon classification Animalia Coleoptera Dytiscidae

1. Exocelina miriae (Balke, 1998) Figs 2, 3, 25

Copelatus (Papuadytes) miriae Balke, 1998: 333; Nilsson 2001: 77 (catalogue).

Papuadytes miriae (Balke, 1998): Nilsson and Fery 2006: 56 (comb. n.).

Exocelina miriae (Balke, 1998): Nilsson 2007: 34 (comb. n.); Toussaint et al. 2014: supplementary figs 1-4, tab. 2.

Type locality.

Papua New Guinea: Morobe Province, Herzog Range, Wagau (Vagau), ca. 06°48'S; 146°48'E, ca. 1300 m a.s.l.

Type material studied.

Paratype: 1 male "Stn. No. 137", "New Guinea: Morobe Dist., Herzog Mts., Vagau, C.4,000ft. 4-17.i.1965", "M. E. Bacchus. B. M. 1965-120", "Paratypus Copelatus miriae sp.n. Balke des. 1997" [red] (NHMW).

Additional material.

Eastern Highlands: 16 males, 15 females "Papua New Guinea: Eastern Highlands, Aiyura, 1670m, 5.iv.2006, 06.21.131S 145.54.398E, Balke & Sagata (PNG 32)" (NHMW, ZSM). 1 male "Papua New Guinea: Eastern Highlands, Aiyura, ditch in forest, 1670 m, 20.v.2006, 06.21.131S 145.54.398E, John & Balke (PNG 69)" (ZSM). 3 males, 1 female "Papua New Guinea: Eastern Highlands, Aiyura, creek, 1670 m, 20.v.2006, 06.21.131S 145.54.398E, John & Balke (PNG 70)" (ZSM). 12 males, 18 females "Papua New Guinea: Eastern Highlands, Onerunka, small creek, red soil /rock, 1700m, 21.v.2006, 06.20.936S 145.46.874E, John & Balke (PNG 71)" (NHMW, ZSM). 5 males, 5 females "Papua New Guinea: Eastern Highlands, Onerunka-Kainantu, 1799m, 14.i.2003, 06 20.561S 145 46.525E, K. Sagata (WB3)" (ZSM). 1 male, 4 females "Papua New Guinea: Eastern Highlands, Yoginofi-Kainantu, 1940m, 14.i.2003, 06 21.483S 145 45.281E, K. Sagata (WB4)" (ZSM). 1 male "390 DNA M Balke", "PNG: EHL, Onerunka-Kainantu, ii.2003, Sagata, DNA M Balke: MB 390" (ZSM). 2 males, 1 female "Papua New Guinea: Eastern Highlands, Abave, small creek, 1500 m, 21.v.2006, 06.17.35S 145.37.681E, John & Balke (PNG 72)" (ZSM). 251 males, 127 females "Papua New Guinea: Eastern Highlands, Kainantu, Yoginofi, 1900m, 9.v.1994, 06.21.799S 145.45.463E, Balke & Sagata (PNG 55)" (NHMW, ZSM). 1 male "385 DNA M Balke", "PNG: EHL, Kainantu, Yoginofi-Kainantu, ii.2003, Sagata, DNA M Balke: MB 385" (ZSM). 20 males, 14 females "Papua New Guinea: Eastern Highlands, Hogu, 1 km E Mt. Barola, 1900m, 9.v.1994, 06.17.556S 145.45.036E, Balke & Sagata (PNG 56)" (NHMW, ZSM). Morobe: 3 females "Stn. No. 139", "New Guinea: Morobe Dist., Herzog Mts., Vagau, C.4,000ft. 4-17.i.1965", "M. E. Bacchus. B. M. 1965-120" (BMNH). 1 male, 3 females "Stn. No. 140A", "New Guinea: Morobe Dist., Herzog Mts., Vagau, C.4,000ft. 4-17.i.1965", "M. E. Bacchus. B. M. 1965-120" (BMNH). 2 males, 1 female "Stn. No. 144", "New Guinea: Morobe Dist., Herzog Mts., Vagau, C.4,000ft. 4-17.i.1965", "M. E. Bacchus. B. M. 1965-120" (BMNH). 1 female "Stn. No. 150", "New Guinea: Morobe Dist., Herzog Mts., Vagau, C.4,000ft. 4-17.i.1965", "M. E. Bacchus. B. M. 1965-120" (BMNH). 42 males, 52 females "Papua New Guinea: Morobe, Wagau, Herzog Mts., 1150m, 19.xi.2006, 06.51.067S 146.48.068E, Balke & Kinibel (PNG 102)", one male with a green label "DNA M.Balke 1380" (NHMW, ZSM). 6 males, 8 females "Papua New Guinea: Morobe, Herzog Mts., 1000m, 20.xi.2006, nr. 06.51.067S 146.48.068E, Balke & Kinibel (PNG 104)" (ZSM). 2 males, 3 females "Papua New Guinea: Morobe, Menyamya, 4-5h towds [towards] Aseki, 1500-2000m, 15.XI.2006, nr 07.14.956S 146.05.687E, Balke & Kinibel, (PNG 100)" (ZSM). 101 males, 54 females "Papua New Guinea: Gulf [sic!], Menyamya, Mt Inji 1700m, 14.xi.2006 nr 07.14.813S 146.01.330E Balke & Kinibel (PNG 96)", one male with a green label "DNA M.Balke 1374" (NHMW, ZSM). 1 female "PAPUA NEW GUINEA Wau, Morobe Prov. Mt. Missim, 1500 m Coldwater Crk. 3 Nov 1985 Col. By MP Kowalski" (ZSM).

Diagnosis.

Beetle medium-sized (TL-H 3.9-4.5 mm); piceous, usually with brownish pronotal sides and head; shiny, with fine but evident punctation and microreticulation; pronotum with distinct lateral bead; male and female antennomere 2 distinctly enlarged, antennomeres 3-6 stout (Fig. 25); protarsomere 4 with large, strongly curved anterolateral hook-like setae; male protarsomere 5 ventrally with anterior band of ca. 60 and posterior row of 13 relatively long, thin setae (Fig. 2A); median lobe evenly curved, with slightly curved, elongate and broadly pointed apex in lateral view, evenly tapering, with rounded apex in ventral view, on both lateral sides with numerous fine setae situated linearly usually on anterior half of distal part of median lobe; paramere without notch, slightly concave on dorsal side and with dense, strong setae on subdistal part and fine proximal setae (Fig. 2 B–D).

Variability.

Specimens from the Eastern Highlands have a shorter apex of the median lobe and more numerous lateral setae situated on almost the whole distal part of the median lobe (Fig. 3).

Distribution.

Papua New Guinea: Morobe and Eastern Highlands Provinces (Fig. 40).