Lobogonodes complicata ( Butler, 1879 )

Wu, Shipher & Chang, Wei-Chun, 2018, Revising the generic characters of Lobogonodes Bastelberger, 1909, with description of a new species from Taiwan (Lepidoptera: Geometridae: Larentiinae), Zootaxa 4433 (3), pp. 434-444 : 440

Lobogonodes complicata ( Butler, 1879)

( Figs 9, 10 , 20 , 28 , 35 )

Cidaria complicata Butler, 1879 , Illustrations of typical Specimens of Lepidoptera Heterocera in the Collection of the British Museum 3: xiv, 57, pl. 55, fig. 4, syntype(s) in BMNH, TL: [Japan], Yokohama.

Microlygris porphyriata complicata: Prout (1914) : 207, pl. 13: a.

Lobogonodes (Microlygris) complicata: Prout (1937) : 105.

Microlygris complicata complicata: Inoue (1982) : 282, 486, pl. 72: 37 (♂), 38 (♀); Wang (1997): 305, fig. syntype; Parsons et al. (1999): 602.

Lobogonodes complicata complicata: Wang (1997) : 305, figs (including the dissected holotype); Choi (2001): 34; Nakajima & Yazaki (2011): 274, pl. 1 – 0 59 – 28, 29.

Lobogonodes complicata: Choi (2012) : 47, figs 32, 112, 192, 271.

Specimens examined. JAPAN. 1♂, Aichi Pref., Iwaya, Minamichita-cho , 19–IV–1987, leg. T. Mano (CHK) ; 1♀, Shizuoka Pref., Kosai City, Shirasuka , 29–VII–1995, leg. S. Iwasaki (CHK).

Diagnosis. This species is very similar to the allopatric L. dactylotypa Prout, 1940 stat. nov. ( Figs 11, 12 , 21 , 29 , 36 , 43 ) but can be distinguished by the more rufous coloration rather than dark brown; the longer saccus; and the wider plate-like signum.

Distribution and bionomics. Distributed in Japan and Korea ( Choi 2012). The records of host plants are Parthenocissus (Vitaceae) ( Sato & Nakajima 1987) including P. tricuspidata ( Nakajima & Yazaki 2011) .

Notes. One syntype is illustrated in Wang (1997, annotated as a holotype in the legend).

Figures

FIGURES 1–15. Habitus of Lobogonodes spp. 1. L. permarmorata (Bastelberger, 1909), Taiwan; 2. Ditto, female, Taiwan; 3. L. shiushioui sp. nov., male, holotype, Taiwan; 4. Ditto, male, paratype, Taiwan; 5. L. erectaria (Leech, 1897), male, Japan; 6. Ditto, female, Japan; 7. L. taiwana (Wileman & South, 1917), female, Taiwan; 8. Ditto; 9. L. complicata (Butler, 1879), male, Japan; 10. Ditto, female, Japan; 11. L. dactylotypa Prout, 1940 stat. rev., male, Taiwan; 12. Ditto, female, Taiwan; 13. L. multistriata (Butler, 1889), male, Japan; 14. Ditto, female, Japan; 15. L. porphyriata (Moore, 1888), male, Nepal. Specimens courtesy of: TFRI (1, 3, 4, 5, 12); ESRI (2, 11), CKY (6, 13–15); CHN (9, 10). Photo: Shipher Wu (1–5, 7–12); Katsumi Yazaki (6, 13–15).

FIGURES 16–23. Male genitalia (without aedeagus) of Lobogonodes spp. 16. L. permarmorata (Bastelberger, 1909), Taiwan; 17. L. shiushioui sp. nov., paratype, Taiwan; 18. L. erectaria (Leech, 1897), Japan; 19. L. taiwana (Wileman & South, 1917), Taiwan; 20. L. complicata (Butler, 1879), Japan; 21. L. dactylotypa (Prout, 1940) stat. rev., Taiwan; 22. L. multistriata (Butler, 1889), Japan; 23. L. porphyriata (Moore, 1888), Nepal. Specimens courtesy of: TFRI (16–19); CHN (20), ESRI (21), CKY (22, 23). Photo: Shipher Wu (16–21); Katsumi Yazaki (22, 23).

FIGURES 24–31. Aedeagus of Lobogonodes spp. 24. L. permarmorata (Bastelberger, 1909), Taiwan; 25. L. shiushioui sp. nov., paratype, Taiwan; 26. L. erectaria (Leech, 1897), Japan; 27. L. taiwana (Wileman & South, 1917), Taiwan; 28. L. complicata (Butler, 1879), Japan; 29. L. dactylotypa Prout, 1940 stat. rev., Taiwan; 30. L. multistriata (Butler, 1889), Japan; 31. L. porphyriata (Moore, 1888), Nepal. Specimens courtesy of: TFRI (24–27); CHN (28), ESRI (29), CKY (30, 31). Photo: Shipher Wu (24–29); Katsumi Yazaki (30, 31).

FIGURES 32–37. Female genitalia of Lobogonodes spp. 32. L. permarmorata (Bastelberger, 1909), Taiwan; 33. L. erectaria (Leech, 1897), Japan; 34. L. taiwana (Wileman & South, 1917), female, Taiwan; 35. L. complicata (Butler, 1879), Japan; 36. L. dactylotypa Prout, 1940 stat. rev., Taiwan; 37. L. multistriata (Butler, 1889), Japan. Specimens courtesy of: TFRI (32, 34, 36); CHN (33, 35), CKY (37). Photo: Shipher Wu (32–36); Katsumi Yazaki (37).

FIGURES 38–43. Life images of Lobogonodes spp. 38. L. permarmorata (Bastelberger, 1909), male, Taiwan; 39. Ditto; 40. L. shiushioui sp. nov., male, paratype, Taiwan; 41. L. erectaria (Leech, 1897), male, Japan; 42. L. taiwana (Wileman & South, 1917), female, Taiwan; 43. L. dactylotypa Prout, 1940 stat. rev., male, Taiwan. Photo: Shipher Wu (38, 41–43); Cheng-Han Ma (39).