Rhyacophila kisszoltani, Kiss, Ottó, 2013

Kiss, Ottó, 2013, Three new species of Rhyacophila (Trichoptera: Rhyacophilidae) from Taiwan and Nepal, Zootaxa 3640 (2), pp. 213-223 : 218-221

Rhyacophila kisszoltani , new species

Figs 13–18

Male. Body length 6.3 mm, length of each forewing 7.5 mm, width of each forewing 3.5 mm, length of each antenna 5 mm. Body, antennae, palpi, wings, abdomen and legs yellowish brown in alcohol.

Male genitalia ( Figs 13–18 ). Segment IX (IX, Figs 13, 15 ) narrow subrectangular in lateral and dorsal views. Segment X (X, Figs 13, 15, 18 ) subquadrate with narrow V-shaped excision on apicodorsal edge in dorsal and caudal views, two parallel vertical margins of segment X with two papillae and with slender process directed ventrocaudad. Apical band (a.b., Figs 13, 18 ) forming vertical and curved horizontal process. Anal sclerites (a.s., Figs 13, 18 ) bulging, paired and separated from each other in caudal view. Tergal strap (t.s., Fig. 18 ) sclerotized with two claws, and serves as a base of the two joining arms of apical band. Endotheca (end., Figs 13–14, 17 ) broad and funnel-like. Distal end of phallicata (ph., Figs 13–14, 17 ) tubular, of uniform diameter and slightly bent dorsocaudad in lateral and ventral views﹔ phallic apparatus without dorsal appendage (d.app.) in lateral view. Retracted parameres (par., Figs 13–14, 17 ) longer than phallicata, with proximal part in sheath, bowl-shaped distal part with hair brush on distal margin in lateral and ventral views. First segment of each inferior appendage (f.s., Figs 16–17 ) broad, moderately long in lateral and ventral views﹔ upper lobe of second joint of each inferior appendage (s.j., Fig. 16–17 ) depressed, dorsal margin undulate, as long as lower lobe with flat, subtriangular process on base of lower margin. Lower lobe ( Fig. 16 ) subtriangular.

Female. Unknown.

Holotype: 3, NEPAL: [Dolakha District], Mt. Kalinchok, 6 km NNE of Muldi (Murre), 27o23’N, 85o58’E, 2835 m elevation, by light trapping, 27 June 1997, leg. Márton Hreblay and Krisztina Csák (gen. prep. No. 115, Ottó Kiss, coll. Mátra Museum).

Paratype: 1 3, EAST-NEPAL: [Taplejung District], Deorali Danda, 1 km N of Yamphudin, 27o23’N, 87o54’E, 1850 m elevation by light trapping, 12 May 1997, leg. Márton Hreblay and Lajos Szécsényi (coll. Ottó Kiss).

Diagnosis. This species belongs to the Rhyacophila scissa Group of the R. philopotamoides Branch ( Schmid 1970) . The male genitalia of the new species are very similar to those of Rhyacophila scissa Morton 1900 [ R. scissa scissa Schmid 1970 ], but differs from them in that:

1. Segment X is subquadrate in dorsal view (not trapezoidal as in R. s. scissa )﹔ with a narrow V-shaped excision (not a broad excision as in R. s. scissa ) on apicodorsal edge﹔ lateral edges straight (not sinuous-convex as in R. s. scissa )﹔ and the two parallel, vertical margins each have two papillae and a slender process directed ventrocaudad (serrated, without papillae or a slender process ventrocaudad in R. s. scissa ).

2. The phallicata is uniform in diameter (the apex is bilobed in R. s. scissa ).

3. The ventral lobe of the phallic apparatus is forked, with two spikes dorsally and one spike laterally (there are only two dorsal spikes in R. s. scissa ).

4. The distal end of each paramere is bowl-shaped (not spatulate as R. s. scissa ) with hair brush on distal margin (not fully covered with hair as in R. s. scissa ).

5. The dorsal margin of the upper lobe of the second joint of each inferior appendage is undulated (not evenly curved as in R. s. scissa ) and is as long as the lower lobe (not shorter than the lower lobe as in R. s. scissa ) and has an obtusely angled, triangular process on the base of its lower margin (without such a subtriangular process in R. s. scissa ). The lower lobe of the second joint of each inferior appendage is subtriangular (not subelliptic as in R. s. scissa ).

Etymology. Named in memory of my father, Zoltán Kiss.

Figures

FIGURES 13 – 18. Rhyacophila kisszoltani new species, holotype male genitalia. 13 L, left lateral view without inferior appendages ﹔ 14 L, phallic apparatus, left lateral view ﹔ 15 D, segments IX and X, dorsal view ﹔ 16 L, right inferior appendage, left mesal view ﹔ 17 V, phallic apparatus, ventral view ﹔ 18 C, vertical distal part of segment X, caudal view. a. b. = apical band ﹔ a. s. = anal sclerite ﹔ end. = endotheca ﹔ f. s. = first segment of paired inferior appendages ﹔ IX. = segment IX ﹔ par. = paired parameres ﹔ ph. = phallicata ﹔ phal. = phallotheca ﹔ r. a. s. = radix of anal sclerite ﹔ s. j. = second joint of paired inferior appendages ﹔ t. b. = tergal band ﹔ ten. = paired tenons of phallotheca ﹔ tend. = tendon of first segment of inferior appendage ﹔ t. s. = tergal strap ﹔ v. l. = ventral lobe of phallicata ﹔ v. p. X. = vertical part of segment X ﹔ X. = segment X.