Tyrtaeus Champion, 1913

Schawaller, Wolfgang & Purchart, Luboš, 2012, Nanocaecus hlavaci gen. & sp. nov. - first record of the tribe Gnathidiini (Coleoptera: Tenebrionidae: Diaperinae) from the Socotra Archipelago, Acta Entomologica Musei Nationalis Pragae 52, pp. 303-314 : 311-312

Tyrtaeus Champion, 1913

Tyrtaeus singularis Grouvelle, 1918 Seychelles (Isl. Mahé)

Gnathidiini incertae sedis

Betschia Dajoz, 1980

Betschia minuta Dajoz, 1980 Madagascar

Remarks. Betschia was established by DAJOZ (1980) as a new genus of the family Colydiidae for Betschia minuta Dajoz, 1980 (by monotypy). IVIE & ŚLIPIŃSKI (1990) transferred the genus Betschia from the family Colydiidae to the Tenebrionidae . Betschia , along with its associated family-group name Betschiini Dajoz,1980 were included under Tenebrionidae incertae sedis in BOUCHARD et al. (2011). The latter authors mentioned that its systematic placement could be close to Gnathidiini . Members of the tribe Gnathidiini possess the tarsal formula 5-5-4 or 4-4-4 (MATHEWS & BOUCHARD 2008) while the genus Betschia has the tarsal formula 3-3-3. Therefore its placement remains uncertain, and for this reason it is considered here as incertae sedis as well.