Tonnoiriella paveli Ježek, 1999

Wagner, Rüdiger & Withers, Phil, 2020, The West-Palearctic species of the genus Tonnoiriella Vaillant, 1971 (Diptera: Psychodidae, Psychodinae), Zootaxa 4728 (2), pp. 183-210 : 189

Tonnoiriella paveli Ježek, 1999

( Figs. 10, 11 )

Tonnoiriella paveli Ježek, 1999 : Časopis Národniho Muzea v Praze Řada přirodovedna 168: 10.

Material: 1♂ holotype, slide mounted: head, antennae (1), palpi (2), thorax (2), wings (2), halter (1), legs (4), ab- domen (1), styli (2), aedeagus complex (1), epandrium and epandrial processes (1); left side white label: Morocco, 8278, High Atlas Mts. , a stream near a wet rocky wall S of Tizi-n-Test approximately 1700 m a.s.l., 2.–3. June 1985, Chvojka leg.; near forest brook with close undergrowth. Right side red label: Tonnoiriella paveli Ježek 1989 ; holo- type Cat. No. 33802, 1996 male . 2♂ (nos 8279, 8280; Cat. Nos. 33803, 33804) with holotype. 1♂ paratype, left side white label: Morocco, 8283, High Atlas Mts. , Toubkai Massif, a streamlet, rocks NW of Lac Ifni (2.500 m a.s.l.), 3.October 1994, Chvojka leg.; right side red label: Tonnoiriella paveli Ježek 1989 , paratype, Cat. No 33807, 1999 male . 1♂ paratype, left side white label: Morocco, 8285; NE of Anto-Atlas Mts, a streamlet of a semidesert area; 1.800 m a.s.l. N of Tizi-n-Bachkoun , 1.June 1985 Chvojka leg. Right side red label: Tonnoiriella paveli Ježek 1989 , paratype, Cat. No 33808, 1999 male (all in collection NMPC) .

Description: Head round, eyebridge of 3 facet rows; distance between eyes 4 facet diameters; postocular bristles present. Palpus segments absolute length: 0.06-0.07 - 0.096 -0.166 mm; relative length: 30-35-48-83. Antenna with scape, pedicel and 14 flagellomeres: absolute length 0.042 -0.054 -0.04-0.04-0.04-0.04-0.04-0.044-0.046-0.044- 0.036-0.034-0.03-0.02 -0.024 - 0.028 mm; relative length 21-27-20-20-20-20-20-22-23-22- 18-17-15-10-12 -14; terminal flagellomere with apiculus. Pairs of digitate ascoids on flagellomeres 5–11.

Wing 3.18 times longer than wide; length 1.84 mm, width 0.58 mm.

Male terminalia ( Fig. 10 ) with hypandrium medially broadened. Gonocoxites are approximately 2.5x longer than wide with a long seta at about middle. Gonostyli about as long as gonocoxites, almost straight. Parameres difficult to identify, small.

Aedeagus with dorsal lobe of the basiphallus terminating in a broad sclerite laterally with elongate spine on one side, and a less sclerotized area on the other side ( Fig. 11 ). Ventral arm bilobed, short lobe connecting at about middle to the lateral side with the distiphallus, long lobe almost straight, tip at about tip of gonocoxite. The ventral distiphallus is a large strongly S-shaped sclerite, margin of baso-lateral end runs parallel to the margin of the dorsal basiphallus lobe; laterally with a tip, in the basal third with a small lobe connecting to the ventral lobe of the basiphallus; the long cross arm almost straight with blunt tips. The dorsal distiphallus has two slightly sinuous lateral lobes with sharp tips; aedeagal sheath with numerous wrinkles.

Epandrium is rectangular; epandrial processes almost straight longer than epandrium with 11 tenacula, length of latter decreasing apically.

Distribution: North Africa, Morocco.

Remarks: T. paveli is a close relative of T. pulchra due to the shape of the ventral distiphallus, bilobed on both sides; the dorsal distiphallus has two large lateral acute tips. The shape of the median part of the hypandrium was not clearly discernable and may differ.

Figures

FIGURES 8–14: 8 T. pulchra, genitalia ventral view; 9 T. pulchra, tip of aedeagus, dorsal view; 10 T. paveli, genitalia ventral view; 11 T. paveli ventral lobe of basiphallus with spineous ventral and dorsal distiphallus sclerites, dorsal view; 12 T. aurasica genitalia ventral view; 13 T. aurasica ventral lobe of basiphallus with spine and flexible area connecting with ventral distiphallus; 14 T. anderssoni, genitalia ventral view (in all figures epandrium, epandrial processes, and proctiger not shown).