Alloraphes yucatani Franz

Jałoszyński, Paweł, 2013, Revision of the Neotropical genus Alloraphes Franz (Coleoptera: Staphylinidae: Scydmaeninae), Zootaxa 3750 (5), pp. 549-568 : 561-562

Alloraphes yucatani Franz

( Figs. 3 , 31–32 , 48–49 , 63 , 68 )

Alloraphes yucatani Franz, 1994: 377 , Fig. 73.

Type material. Holotype: Mexico (Campeche): ♂, three labels ( Fig. 63 ): "Umg. Camp[crossed out "a"]eche / Yucatan, Mexico " with "Mx8" on reverse side [white, printed; reverse handwritten in blue ink]; " Alloraphes / yucatanensis [sic!] m. / det. H. Franz" [white, handwritten and printed]; " Holotypus " [red, handwritten] (NHMW). Paratypes (1 ♂, 2 ♀): same data as holotype, except for yellow " paratypus " label; one female without locality label (NHMW).

Revised diagnosis. Male: frons in anterior part flat, in posterior part with transverse impression sharply demarcated from convex vertex; pronotum with nearly evenly arcuate ante-basal transverse groove with distinct median pit; elytra modified, each with strongly impressed apical region slightly protruding caudad; aedeagus in ventral view with distinctly asymmetrical apex of median lobe, long and flat apical projections and rounded apices of parameres each bearing three thick and long setae. Female: antennomere XI barely longer than wide; antennomeres III–X as long as broad or slightly transverse; frons between antennal insertions with flat substriangular area sharply delimited from convex posterior part of head.

Redescription. Body of male ( Fig. 3 ) moderately strongly convex, brown, covered with light brown vestiture; BL 0.90–0.96 mm.

Head broadest at large, strongly convex and moderately coarsely faceted eyes, HL 0.13 mm, HW 0.19 mm; tempora absent, eyes adjacent to occipital constriction; vertex distinctly convex, its anterior margin sharply demarcated from transverse arcuate posterior frontal impression; anterior part of frons flat; supraantennal tubercles feebly marked. Punctures on frons and vertex inconspicuous, fine; setae short, sparse and suberect. Antennae slender, AnL 0.35 mm; antennomere I about twice as long as broad; II 1.8x as long as broad; III barely noticeably transverse; IV–VI each slightly but distinctly transverse; VII–X strongly transverse; XI only about 1.3x as long as broad and about 1.5x as long as X.

Pronotum about as long as broad, broadest in anterior fourth; PL 0.23–0.26 mm, PW 0.23–0.25 mm. Anterior margin and lateral margins in anterior third strongly rounded; sides distinctly but slightly narrowing caudad, pronotum indistinctly constricted near posterior fourth; posterior margin slightly arcuate; ante-basal transverse groove distinct and nearly evenly arcuate, with small median pit and large but shallow lateral impressions. Punctures on pronotal disc inconspicuous, fine and sparse; setae moderately dense, short and nearly recumbent, sides of pronotum with moderately long and disordered bristles.

Elytra more convex than pronotum, oval with slightly protruding apices, broadest slightly anterior to middle; EL 0.55–0.58 mm, EW 0.40 mm, EI 1.38–1.44; sub-humeral lines sharply marked and slightly shorter than 1/3 EL; elytral apices spoon-shaped, each separately protruding posteriorly and strongly impressed. Punctures on elytra distinct but small and shallow, separated by spaces 1–1.5x as wide as puncture diameters; setae moderately long, sparse and only slightly suberect. Hind wings well developed.

Legs long and slender; all tibiae straight.

Aedeagus ( Figs. 31–32 , 48–49 ) slender and lightly scleroitized; AeL 0.23 mm; in ventral view apical part of median lobe distinctly asymmetrical and with deeply but narrowly emarginated apical margin; apical projections long and flattened dorso-ventrally; parameres in lateral view broad and slightly narrowing in apical parts, apices rounded, each paramere with three thick and long setae.

Female. Similar to male but with distinctly smaller eyes, indistinct posterior transverse impression on frons and unmodified elytral apices. BL 0.90 mm; HL 0.13 mm, HW 0.20 mm, AnL 0.35 mm; PL 0.23 mm, PW 0.23 mm; EL 0.55 mm, EW 0.40 mm, EI 1.38.

Distribution. Southeastern Mexico (Yucatán) ( Fig. 68 c).

Remarks. Franz (1994) gives additional collecting data: the holotype male was taken by sifting dry leaf litter in a forest not far from Edzna on 4.vi.1992, while the paratype male and female were collected by sifting dry leaf litter in Dzibilchaltún (spelled as Szibilchaltun) near Mérida on 9.vi.1992 (both localities are Mayan archaeological sites).

Figures

FIGURE 68. Distribution of Alloraphes: a, A. jamaicae Franz; b, A. chiapasensis Franz; c, A. yucatani Franz; d, A. lenkoi Franz; e, A. peruanus Franz; f, A. dentatus sp. n.; g, A. peckorum sp. n.; h, A. myrmecophilus Franz stat. rest.; i, A. bolivarensis Franz; j,? Alloraphes magnus Franz.

FIGURES 1 – 12. Dorsal (1 – 9, 11 – 12) and laterodorsal (10) habitus of Alloraphes Franz. Alloraphes jamaicae Franz, holotype male (1); A. chiapasensis Franz, holotype male (2); A. yucatani Franz, holotype male (3); A. lenkoi Franz, paratype male (4) and paratype female (5); A. peruanus Franz, holotype male (6) and paratype female (7; but see the Remarks section in the redescription); A. dentatus sp. n., holotype male (8); A. peckorum sp. n., holotype male (9 – 10); A. myrmecophilus Franz, stat. rest., paratype male (11);? Alloraphes magnus Franz, holotype female (12).

FIGURES 27 – 43. Aedeagus (27 – 34, 36 – 43) and female genitalia (35) of Alloraphes Franz in ventral (27, 29, 31, 33, 35, 36, 38, 40, 42) and lateral (28, 30, 32, 34, 37, 39, 41, 43) views. Alloraphes jamaicae Franz, holotype (27 – 28); A. chiapasensis Franz, holotype, parameres missing (29 – 30); A. yucatani Franz, holotype (31 – 32); A. lenkoi Franz, paratypes (33 – 35); A. peruanus Franz, holotype (36 – 37); A. dentatus sp. n., holotype (38 – 39); A. peckorum sp. n., holotype (40 – 41); A. myrmecophilus Franz, stat. rest., paratype (42 – 43). Abbreviations: ap, apical projection; bpa, basal pumping apparatus; ov, egg; sp, spermatheca.

FIGURES 44 – 60. Aedeagus (44 – 51, 53 – 60) and female genitalia (52) of Alloraphes Franz in ventral (44, 46, 48, 50, 52, 53, 55, 57, 59) and lateral (45, 47, 49, 51, 54, 56, 58, 60) views. Alloraphes jamaicae Franz, holotype (44 – 45); A. chiapasensis Franz, holotype, paramere detached (46 – 47); A. yucatani Franz, holotype (48 – 49); A. lenkoi Franz, paratypes (50 – 52); A. peruanus Franz, holotype (53 – 54); A. dentatus sp. n., holotype (55 – 56); A. peckorum sp. n., holotype (57 – 58); A. myrmecophilus Franz, stat. rest., paratype (59 – 60). Abbreviations: ap, apical projection; bpa, basal pumping apparatus; ov, egg; sp, spermatheca.

FIGURES 61 – 67. Original sets of labels pinned under historical type specimens. Alloraphes jamaicae Franz, holotype (61); A. chiapasensis Franz, holotype (62); A. yucatani Franz, holotype (63); A. lenkoi Franz, paratype (64); A. peruanus Franz, holotype (65); A. myrmecophilus Franz, stat. rest., paratype (66);? Alloraphes magnus Franz, holotype (67).