Alloraphes jamaicae Franz

Jałoszyński, Paweł, 2013, Revision of the Neotropical genus Alloraphes Franz (Coleoptera: Staphylinidae: Scydmaeninae), Zootaxa 3750 (5), pp. 549-568 : 555-558

Alloraphes jamaicae Franz

( Figs. 1 , 22–24 , 27–28 , 44–45 , 61 , 68 )

Alloraphes jamaicae Franz, 1980a: 213 , Figs. 199a, b.

Type material. Holotype: Jamaica: ♂, three labels ( Fig. 61 ): "Blue Mountains / Jamaica, lg. Franz" with "SA120" on reverse side [white, printed; reverse handwritten in blue ink]; " Alloraphes / jamaicae m. / det. H. Franz" [white, handwritten and printed]; " Typus " [red, handwritten] (NHMW). Paratype: 1 ex., same data as holotype, except for yellow " paratypus " label and "SA121" on reverse side of locality label (NHMW).

Revised diagnosis. Male: frons with narrow and sharply demarcated transverse impression between eyes; pronotum with arcuate ante-basal transverse groove, without median pit; elytra modified, with distinctly impressed apical region; aedeagus in ventral view with distinctly asymmetrical apex of median lobe, short apical projections and broad, irregular in shape apices of parameres each bearing three thick and long setae. Diagnostic characters of females unknown.

Redescription. Body of male ( Fig. 1 ) moderately strongly convex, brown, covered with light brown vestiture; BL 0.95 mm.

Head ( Fig. 22 ) broadest at large, strongly convex and moderately coarsely faceted eyes, HL 0.13 mm, HW 0.20 mm; tempora in dorsal view rudimentary; vertex distinctly convex; frons flattened and with deep transverse impression between eyes (lateral margins of impression distant from mesal margins of eyes); supraantennal tubercles feebly marked. Punctures on frons and vertex inconspicuous, fine; setae short, sparse and suberect. Antennae slender, AnL 0.43 mm; antennomeres I and II each about twice as long as broad; III slightly longer than broad; VI about 1.2x as long as broad; V–VII each about as long as broad; VIII slightly transverse; IX–X each distinctly transverse; XI about 1.5x as long as X and 1.5x as long as broad.

Pronotum ( Fig. 23 ) about as long as broad, broadest in anterior third; PL 0.25 mm, PW 0.24 mm. Anterior margin and lateral margins in anterior half strongly rounded; sides distinctly narrowing caudad, pronotum distinctly constricted near posterior fourth; posterior margin slightly arcuate; ante-basal transverse groove distinct and arcuate, without median pit and with shallow lateral pits. Punctures on pronotal disc inconspicuous, fine and sparse; setae sparse, short and suberect; sides of pronotum with long and disordered bristles.

Elytra more convex than pronotum, oval, broadest near anterior third; EL 0.58 mm, EW 0.43 mm, EI 1.35; sub-humeral lines moderately distinct, as long as 1/4 EL; elytral apices distinctly impressed except for elytral suture which remains raised. Punctures on elytra distinct but small and shallow, separated by spaces 1–1.5x as wide as puncture diameters; setae moderately long, sparse and suberect. Hind wings well developed.

Legs long and slender; all tibiae straight.

Aedeagus ( Figs. 27–28 , 44–45 ) slender and moderately darkly sclerotized; AeL 0.23 mm; in ventral view apical part of median lobe distinctly asymmetrical and with deeply but narrowly emarginated apical margin; apical projections short and obscured by overlapping apices of parameres; parameres in lateral view with strongly expanded apical parts, in ventral view irregular in shape, each with three thick and long setae.

Female. Unknown.

Distribution. Greater Antilles: eastern part of Jamaica ( Fig. 68 a).

Remarks. The type series of A. jamaicae consists of a well-preserved holotype male and remnants of a paratype, with the head, prothorax and the right elytron missing. According to Franz (19980) it is a male. However, the holotype male has a distinctly modified apical region of elytra, while the paratype has the elytron unmodified. It is either a male of a different species or a female. This specimen was not dissected to avoid further damages. Franz (1980a) gives more precise collecting data: Hardwar-Gap, 1500 m alt., from rotten trunks and stumps, 20.ix.1968.

Figures

FIGURE 68. Distribution of Alloraphes: a, A. jamaicae Franz; b, A. chiapasensis Franz; c, A. yucatani Franz; d, A. lenkoi Franz; e, A. peruanus Franz; f, A. dentatus sp. n.; g, A. peckorum sp. n.; h, A. myrmecophilus Franz stat. rest.; i, A. bolivarensis Franz; j,? Alloraphes magnus Franz.

FIGURES 1 – 12. Dorsal (1 – 9, 11 – 12) and laterodorsal (10) habitus of Alloraphes Franz. Alloraphes jamaicae Franz, holotype male (1); A. chiapasensis Franz, holotype male (2); A. yucatani Franz, holotype male (3); A. lenkoi Franz, paratype male (4) and paratype female (5); A. peruanus Franz, holotype male (6) and paratype female (7; but see the Remarks section in the redescription); A. dentatus sp. n., holotype male (8); A. peckorum sp. n., holotype male (9 – 10); A. myrmecophilus Franz, stat. rest., paratype male (11);? Alloraphes magnus Franz, holotype female (12).

FIGURES 22 – 26. Alloraphes jamaicae Franz, holotype male (22 – 24); A. myrmecophilus Franz stat. rest., paratype male (25);? Alloraphes magnus Franz, holotype female (26). Head in ventral view (22); prothorax in ventral view (23); pterothorax in ventral view (24); body venter (25); and mesothorax in ventral view (26). Abbreviations: aest 2, mesanepisternum; aest 3, metanepisternum; ai, asetose impression; amc, anterior marginal carina of mesocoxal cavity; ar, anterior ridge; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; cx, coxa; el, elytron; epm 3, metepimeron; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; lss, lateral suture of submentum; mc, median carina of mesoventrite; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mesostipes; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; pl, posterior lobe of mesocoxal projection; pre, prepectus; ppf, palpifer; psp, prosternal process; submentum; tm, temple; v 3, metaventrite

FIGURES 27 – 43. Aedeagus (27 – 34, 36 – 43) and female genitalia (35) of Alloraphes Franz in ventral (27, 29, 31, 33, 35, 36, 38, 40, 42) and lateral (28, 30, 32, 34, 37, 39, 41, 43) views. Alloraphes jamaicae Franz, holotype (27 – 28); A. chiapasensis Franz, holotype, parameres missing (29 – 30); A. yucatani Franz, holotype (31 – 32); A. lenkoi Franz, paratypes (33 – 35); A. peruanus Franz, holotype (36 – 37); A. dentatus sp. n., holotype (38 – 39); A. peckorum sp. n., holotype (40 – 41); A. myrmecophilus Franz, stat. rest., paratype (42 – 43). Abbreviations: ap, apical projection; bpa, basal pumping apparatus; ov, egg; sp, spermatheca.

FIGURES 44 – 60. Aedeagus (44 – 51, 53 – 60) and female genitalia (52) of Alloraphes Franz in ventral (44, 46, 48, 50, 52, 53, 55, 57, 59) and lateral (45, 47, 49, 51, 54, 56, 58, 60) views. Alloraphes jamaicae Franz, holotype (44 – 45); A. chiapasensis Franz, holotype, paramere detached (46 – 47); A. yucatani Franz, holotype (48 – 49); A. lenkoi Franz, paratypes (50 – 52); A. peruanus Franz, holotype (53 – 54); A. dentatus sp. n., holotype (55 – 56); A. peckorum sp. n., holotype (57 – 58); A. myrmecophilus Franz, stat. rest., paratype (59 – 60). Abbreviations: ap, apical projection; bpa, basal pumping apparatus; ov, egg; sp, spermatheca.

FIGURES 61 – 67. Original sets of labels pinned under historical type specimens. Alloraphes jamaicae Franz, holotype (61); A. chiapasensis Franz, holotype (62); A. yucatani Franz, holotype (63); A. lenkoi Franz, paratype (64); A. peruanus Franz, holotype (65); A. myrmecophilus Franz, stat. rest., paratype (66);? Alloraphes magnus Franz, holotype (67).