Coccoidea

Hodgson, Chris, 2005, The adult males of Coccoidea (Hemiptera: Sternorrhyncha) known from Greenland, Zootaxa 907, pp. 1-48 : 2-4

KEY TO ADULT MALE COCCOIDEA KNOWN FROM GREENLAND *

(for an explanation of terms used, see the Figures)

1. Penial sheath at least 4x longer than wide; scutum with a large quadrangular membranous area medially; ocular sclerites and genae covered in polygonal reticulations; loculate pores entirely absent (apart from in glandular pouches); body with frequent fleshy setae, even on abdomen; hamulohalteres absent ................................................. ................................. COCCIDAE ( Pulvinaria glacialis Gertsson & Hodgson ) (Fig. 9)

­ Penial sheath never more than 2x longer than broad; scutum without a large quadrangular area medially; polygonal reticulations absent from genae and restricted to around each simple eye on ocular sclerites; loculate pores usually present, at least on head but also generally on abdomen; body without fleshy setae on abdomen; hamulohalteres present...........................................................................................................................2

2. Postoccipital ridge strong; scutum with two small membranous areas medially; scutellum with a well­developed scutellar ridge; claw digitules capitate and longer than claw; loculate pores absent apart from on head; apical antennal segment almost as wide as long; cranial apophysis well developed and trilobate; ostioles absent............... ................................................................................ ERIOCOCCIDAE ( Figs 7 & 8 ).....8

­ Postoccipital ridge weak or absent; scutum without two small membranous areas medially; scutellum without a scutellar ridge; claw digitules setose and much shorter than claw; loculate pores often frequent and nearly always present in pleural region of

abdomen; apical antennal segment at least twice as long as broad; cranial apophysis poorly developed or absent; ostioles present........ PSEUDOCOCCIDAE (Figs 1–6)...3 3. Apterous; mesothorax mainly membranous; postoccipital ridge entirely absent .......... ............................................... Chorizococcus multiporus Gertsson & Hodgson ( Fig. 2 )

­ Macropterous; mesothorax heavily sclerotised; postoccipital ridge present................. 4

4. With two pairs of glandular pouches, one pair on abdominal segment VII and the other on segment VIII; fleshy setae on antennae setose and at least twice as long as width of flagellar antennal segments ................................ Peliococcus balteatus (Green) (Fig. 3)

­ With only one pair of glandular pouches, on abdominal segment VIII; fleshy setae on antennae thick, with a blunt apex, clearly different from hair­like setae and each about as long as width of flagellar antennal segments............................................................ 5

5. Several postmesospiracular setae present anterior to basisternum; fleshy setae clearly differentiated from hair­like setae on head; each gena with 4+ genal setae, some often fleshy; small convex pores on ventral epicranium of head .......................................... 6

­ Postmesospiracular setae very few or absent anterior to basisternum; fleshy setae not differentiated from hair­like setae on head; each gena with 3 or fewer genal setae, none fleshy; pores on head, when present, on dorsal epicranium.......................................... 7

6. Fleshy setae on head quite short; 3rd, 4th and 5th antennal segments all equal to or shorter than 2nd segment; postmesospiracular disc­pores and setae few immediately anterior to basisternum; fleshy setae of antennae short (17–20µm on subapical segment), about width of antennal segments ....................................................................... ........................................................... Trionymus bocheri Gertsson & Hodgson ( Fig. 4 )

­ Fleshy setae on head almost as long as hair­like setae; 3rd antennal segment longer than 2nd segment and 3rd and 4th segments subequal in length; postmesospiracular disc­pores few or absent but setae quite abundant immediately anterior to basisternum; fleshy setae of antennae long (30µm on subapical segment), longer than width of antennal segments ............................. Trionymus elymus Gertsson & Hodgson ( Fig. 5 )

7. With 0 or 1 loculate pore on dorsal surface of head; pleural region of each abdominal segment with 0 or 1 loculate pore; abdominal segment VIII with a pair of transverse sclerotisations mediolaterally on sternite; loculate pores absent from dorsal and ventral surfaces of abdomen; with several metatergal setae ...................................................... .............................................. Atrococcus groenlandensis Gertsson & Hodgson (Fig. 1)

­ With several loculate pores on dorsal surface of head; pleural region of each abdominal segment with 2–5+ loculate pores; abdominal segment VIII without a pair of transverse sclerotisations mediolaterally on sternite; loculate pores present on dorsal and ventral surfaces of abdomen; with only 1 pair of metatergal setae ................................. ................................................................................. Trionymus thulensis Green ( Fig. 6 )

8. Length of fleshy setae on antennae equal to or less than width of flagellar segments of antennae; fleshy setae with a blunt apex; all pores on head quite small ........................ ......................................................................... Eriococcus pseudinsignis Green ( Fig. 8 )

­ Length of fleshy setae on antennae clearly greater than width of flagellar segments of antennae; fleshy setae with sharply rounded apices; some pores on head quite large ... ........................................................................ Eriococcus munroi (Boratynski) ( Fig. 7 )

Figures

FIGURE 2. Adult male of Chorizococcus multiporus Gertsson & Hodgson. Where G = multilocular disc­pores; H = minute pore; J = apical antennal segment; K = latero­ventral view of genital segment; M = lateral sclerotisation.

FIGURE 4. Adult male of Trionymus bocheri Gertsson & Hodgson. Where A = cranial apophysis; D = convex simple pore on head; E = fleshy seta; G = multilocular disc­pores; H = minute pore; J = apical antennal segment; K = dorsal and ventral view of genital segment. And where astn 1 s = anteprosternal setae; caps = capitate setae; fs = fleshy setae; hs = hair­like setae; lps = lateral prothoracic setae; lpns = lateral pronotal setae; mpns = medial pronotal setae; pts = post­tergital setae, and stn 1 s = prosternal setae. For most other abbreviations see Figs. 1 & 3.

FIGURE 5. Adult male of Trionymus elymus Gertsson & Hodgson. Where A = cranial apophysis; D = convex simple pore on head; G = multilocular disc­pores; H = minute pore; J = apical antennal segment; K = lateral view of genital segment; L = tarsus plus claw.

FIGURE 6. Adult male of Trionymus thulensis Green. Where A = cranial apophysis; E = fleshy setae; G = multilocular disc­pores; H = minute pore; J = apical antennal segment; K = lateral view of genital segment; L = tarsus plus claw.

FIGURE 7. Adult male of Eriococcus munroi (Boratynski). Where A = cranial apophysis; D = dorsal pores on head; E = fleshy seta; F = hair­like seta; J = apical antennal segment; K = lateroventral view of genital segment; L = tarsus + claw, and M = campaniform pore on trochanter. And where an = anus; cd = claw denticle; cdt = claw digitules; fs = fleshy setae; hs = hair­like setae; por = postoccipital ridge; sclr = scutellar ridge, and tibs = large spur­like setae on tibia.

FIGURE 8. Adult male of Eriococcus pseudinsignis Green. Where A = cranial apophysis; D = dorsal pore on head; E = fleshy seta; K = dorsal and ventral view of genital segment, and L = tarsus + claw.