Bembidion (Plataphus) pseudolucillum Netolitzky, 1938

Neri, Paolo & Toledano, Luca, 2021, Geographic and taxonomic notes, addenda and corrigenda on the subtribe Bembidiina Stephens, 1827 of the 2017 ' Catalogue of Palaearctic Coleoptera' (Coleoptera, Carabidae, Bembidiina), ZooKeys 1044, pp. 563-587 : 563

Bembidion (Plataphus) pseudolucillum Netolitzky, 1938 comb. nov. Figures 2 , 9

Bembidion (Peryphus) pseudolucillum Netolitzky, 1938

Material examined.

1 ♀, " Japan / Settsu / Katsuoji / 16.VI.24 / J.E.A. Lewis [printed] // 199 [handwritten] // Type [red, printed] // pseudolucillum Net. / Type Netolitzky 37 // Peryphus // H.E. Andrewes coll. / B.M. 1945-97 [printed]" (NHMUK) . 3 ♀♀, on the same pin, "Oku Nikko / Nabagawa / 8.9.1937 / Jano, Jap. [handwritten] // pseudolucillum / det. Netolitzky [handwritten] // H.E. Andrewes coll. / B.M. 1945-97 [printed]" (NHMUK) . 1 ♀, " Japan / Settsu / Katsuoji / J.E.A. Lewis [handwritten] // [one deleted label] // Peryphus sp. / striis mediocr- / punctat [illegible] / dt Netolitzky [handwritten] // mit Type / Brit. Museum / identisch Net. [handwritten] // Coll. / Netolitzky [printed] // pseudolucillum Net / Cotypus Netolitzky 37 [handwritten] // CO / TYPUS [red, printed] // Coll. / Netolitzky [printed]" (NHMW) . 1 ♂, "Oku Nikko / Nabagawa / 8.9.37 Jano [handwritten] // pseudolucillum / dt. Netolitzky [handwritten, contoured in red] // coll. / Netolitzky [printed]" (NHMW); from the handwritten label contoured in red we deduce that this last specimen was considered by Netolitzky as a paratype . 1 ♂, 2 ♀♀, on the same pin, "Oku Nikko / Nabagawa / 8.9.37 Jano [handwritten] // coll. / Netolitzky [printed] // pseudolucillum / dt. Netolitzky [handwritten]" (NHMW); we mounted the ♂ on a separate pin, the labels have been photocopied in order to mount the specimens on separate pins, and the ♂, dissected, bears the slide with the aedeagus mounted in euparal on the same pin. 1 ♂, " Kibune , Kyoto / 12-IV-1950 / Leg. S. Uéno [handwritten] // Acqu.-Nr. / 1951-23 [printed] // Coll. / Meyer // pseudolucillum / Netolitzky 1938 [handwritten] / Det. S. Uéno [printed]" (NHMW) (Fig. 2 ) . 2 ♀♀, " Kibune , Kyoto / 12-IV-1950 / Leg. S. Uéno [handwritten] // Acqu.-Nr. / 1951-23 [printed] // Coll. / Meyer // pseudolucillum [handwritten] / det. Paul Meyer [printed]" (NHMW) . 1 ♂, " Japan G. Lewis // Japan. Lewis // Sharp Coll. 1905-313 // Bembidion cfr. pseudolucillum det. L. Toledano 2008 // CTVR Coll. Toledano Verona - Italy " (CTVR) . 1 ♂, "Kujukawa / Aomori Pref. / Honehu Japan / 10.9.54 [handwritten] // Bembidion / pseudolucillum / Netolitzky 1938 / det. L. Toledano 2011 [printed] // CTVR / Coll. Toledano / Verona - Italy [printed]" (CTVR) . 7 ♀♀, "Numata, Yumma Pref., Honshu Japan " (CTVR, PN) . 1 ♀, "Amagi-Pass, Kawazu-cho, Shizuoka pref. " (CTVR) . 1 ♀, "Sanjo-valley, Tabayama vill., Yamanashi-pref. " (CTVR) . 1 ♀, " J. Honshu, Shiga-ken, Mikunidakeyama, E slope, 700 m " (CTVR) . 2 ♀♀, 3 ♂♂, " Japan, Mt. Hiba , Saljo, Hiroshima pref. " (CTVR, PN) .

Notes.

Netolitzky (1938) describes Bembidion (Peryphus) pseudolucillum from Settsu (Japan), assigning it to the group of B. nitidulum Marsham, 1802, s. l. (= B. deletum Serville, 1821); again Netolitzky (1943a, 1943b) found it difficult to find a correct position for the species in his Bestimmungs-Tabellen: he mentions similarities both with species of two subgenera, Bembidionetolitzya Strand, 1929 and Peryphus Dejean, 1821, but preferred the latter. Jedlička (1965), in his monograph on the Bembidiini of Eastern Asia, includes the species in the keys for subgenus Plataphus Motschulsky, 1864 and that of the subgenus Plataphus Peryphus . The species is listed in "incertae sedis" in the last catalogues (e.g., Marggi et al. 2003; Lorenz 2005; Marggi et al. 2017).

Conclusions.

The examination of the holotype and of the aedeagus (Fig. 9 ) shows that the species belongs to subgenus Bembidion Plataphus Motschulsky, 1864. The doubts regarding the systematic position of the species are due to its pronotal convexity, more evident than in other species of the subgenus. We added to all the non-type specimens the following label: Bembidion (Plataphus) pseudolucillum Net. - det. Neri and Toledano 2020. The species length varies from 3.6 to 4.7 mm, the aedeagus from 0.75 to 0.84 mm.

Figures

Figures 1 - 4. Habitus of 1 Bembidion (Peryphanes) sanatum Bates, 1883, holotype (NHMUK), 5.60 mm 2 B. (Plataphus) pseudolucillum Netolitzky, 1938 from Kibune, Kyoto (NHMW), 4.20 mm 3 B. (Peryphus) psuchrum Andrewes, 1935 Inde, Himachal Pradesh, Mahari, 3500 m (CTVR), 3.50 mm 4 labels of the holotype of Bembidion sapporense Jedlicka (NMPC). Photographs: Luca Toledano.

Figures 5 - 9. Median lobe of the aedeagi of 5 Bembidion (Peryphanes) sanatum Bates, 1883, holotype (NHMUK), 1.55 mm 6 B. (P.) dostali Kirschenhofer, 1984, paratype (NHMW), 1.53 mm 7 B. (P.) lulinense Habu, 1973 (drawing from Habu 1973), 1.41 mm 8 Bembidion (Peryphus) psuchrum Andrewes, 1935, holotype (NHMUK), 0.91 mm 9 Bembidion (Plataphus) pseudolucillum Netolitzky, paratype (NHMW), 0.80 mm.