Teuloma Volynkin & N. Singh, 2019

Singh, Navneet, Kirti, Jagbir Singh, Datta, Harvinder Singh, Joshi, Rahul & Volynkin, Anton V., 2019, A review of the genus Dolgoma Moore from India, with notes on the genus taxonomy and descriptions of a new genus, four new species and a new subspecies (Lepidoptera, Erebidae, Arctiinae, Lithosiini), Zootaxa 4683 (1), pp. 33-54 : 40

Genus Teuloma Volynkin & N. Singh , gen. nov.

urn:lsid:zoobank.org:act:57E340A4-9F23-4762-A6CA-7A6D77E01764

Type species: Lithosia nebulosa Walker, 1862 .

Diagnosis. Morphologically, the new genus has a transitional position between Teulisna and Dolgoma . The genus is characterized by two autapomorphic features: (1) a bilobate apex of distal saccular process: one lobe is plate-like, weakly setosed and directed dorsally, while the second one is curved dorsally, twisted and densely covered with strong setae (marked on Fig. 48 ) (that is unilobate in both, Teulisna and Dolgoma ); (2) the presence of a rugose tubelike dorsal extension of juxta heavily connected to aedeagus (marked on Fig. 43 ). In addition, compared to Teulisna , the genus Teuloma has no scent scale modifications to the male forewing and extreme distortion to the venation as well. The male genitalia of Teuloma species are very uniform and differ from those of Teulisna (illustrated by Holloway 2001, fig. 67) by the broader juxta with a deep medial apical concavity and the absence of coremata in vinculum. Compared to those of Dolgoma , the male genitalia of the new genus have the uncus being narrow and laterally flattened (that is much broader and volumetric in Dolgoma ), the scaphium being more heavily sclerotized and sticklike (that is weakly setose in Dolgoma ), the rectangular and weakly sclerotized vinculum having thin distal margin (in Dolgoma , the vinculum is more or less U-shaped and has more heavily sclerotized margins), the longer distal saccular process and lack spinules in vesica. The juxta shape of Teuloma also resembles that of Wittia (illustrated by Dubatolov & Zolotuhin 2011 and Witt et al. 2011) due to the presence of short dentate processes. However, in Wittia these processes are transtilla ones fused with juxta (the genitalia structure of Wittia sororcula is discussed by Witt et al. (2011)), while in Teuloma transtillae have no processes and are not fused with juxta. In addition, the shape of distal saccular process of Teuloma is principally different from that of Wittia , which is simple, elongate and strongly curved dorsad. The female genitalia of Teuloma are characterized by the corpus bursae situated laterally, having a narrow neck-like posterior section, and a large and twisted appendix bursae having very broad basal section.

Remark. The male genitalia of Wittia klapperichi ( Daniel, 1954) are similar to those of Teuloma due to the shape of the distal saccular process. However, the presence of a dentate medial lobe of valva and a transtillar process make impossible its association with Teuloma . The generic placement of klapperichi needs further clarification. Here we provisionally leave this species under the genus Wittia .

Etymology. The new genus name is a combination of the generic names Teulisna and Dolgoma .

Figures

FIGURES 45–48. Teuloma spp., male genitalia. 45, T. falsinebulosa, holotype, NE India, slide NHMUK010315252 Volynkin (©NHMUK); 46, T. falsinebulosa, paratype, NE India, prep. H. S. Datta; 47, T. nebulosa, NE India, prep. H. S. Datta; 485, T. nebulosa, S Laos, slide AV4832 Volynkin.

FIGURES 41–44. Teuloma spp., male genitalia. 41, T. microoblitterans, holotype, NE India, prep. Joshi; 42, T. microoblitterans, paratype, NE India, prep. H. S. Datta; 43, T. neooblitterans, holotype, NE India, prep. H. S. Datta; 44, T. montanebula, SW Thailand, slide AV4694 Volynkin.