Rubus candicans Weihe ex Reichenbach (1832: 601)

Akopian, Janna, 2024, Taxonomic synopsis of the genus Rubus (Rosaceae) in Armenia, Phytotaxa 677 (2), pp. 125-143 : 131

5. Rubus candicans Weihe ex Reichenbach (1832: 601) . Type:— GERMANY. Sachsen, s.d, s.c., s.n (W). ( Fig. 3 C, D )

= R. thyrsoideus Wimmer subsp. candicans (Weihe) Focke (1902: 485) .

Distribution in Armenia: Ijevan floristic region. In shrubberies of steppe zone, broad-leaved forests, river-bed forests, alongside rivers and brook banks, along garden hedges, at an altitude of 700–1400 m above sea level.

Phenology: Flowering June–August; fruiting September–October.

General distribution: Caucasus (Great Caucasus, Transcaucasia), North and Middle Europe, Crimea.

Selected specimens. ARMENIA: Noyemberyan district, Noyemberyan city, along the fences, 25 June 1954, Mulkidzhanyan 66877 ( ERE) ; Akhtala , Debet river valley, 25 June 1954, M ulkidzhania n 63240 ( ERE) ; Alaverdi region , village Ayrum × Mets Ayrum, along the road, 25 June 1954, Mulkidzhanyan 63241 ( ERE) ; Akhsu × Tsakhkavan, on the edge of an oak-hornbeam forest 900–1200 m above sea level, 24 June 1954, Mulkidzhanyan 66774 ( ERE) ; Alaverdi region , Ayrum village × Haghpat station, Debet river valley, 9 August 1954, Mulkidzhanyan 63243 ( ERE) ; Noyemberyan district , vicinity of Ayrum station, right side of the Debet River, oak and hornbeam forest, 22 April 1955, Karapetyan, Aslanyan 58626 ( ERE) ; Shamshadin district , Aygedzor × Kolagiran, 23 September 1960, Mulkidzhanyan 67705 ( ERE) .

Series 3. Hedycarpi Focke (1877: 190).

Annual shoots arcuately curved, scattered pubescent or (almost) glabrous; leaves 5-foliolate, somewhat leathery, unevenly toothed, shortly pointed. Inflorescence narrowed upwards, strong, many-flowered.

6. Rubus ibericus Yuzepchuk (1924 –1925: 153); Yuzepchuk (1941: 26). Lectotype (designated by Kutateladze: 1971):— GEORGIA. Somchetia, inter opp. Tiflis et pag. Kodzhory ad viam, 30 July 1923, S. Juzepczuk 12 (TBI). ( Fig. 3 E, F ).

Distribution in Armenia: Zangezur floristic region. Alongside river banks, roads, at the altitude of 1200–1500 m above sea level.

Phenology: Flowering June–August; fruiting September–October.

For the flora of Armenia R. ibericus is reported for the first time.

General distribution: East and South Transcaucasia, Talysh, East Anatolia.

Selected specimens. ARMENIA: Zangezur, outskirts of Goris . 4 September 1949, Karapetyan 135970 ( ERE) ; Syunik province, Kapan district , N Kapan, road to Yegheg, c. 2.6 km from village, 1289 m, 39 o 16'43''N / 46 o 25'55''E, 21 June 2004, Vitek, Tamanyan, Fayvush, Oganesyan, Ter-Voskanyan 202283 ( ERE) ; Syunik province, c. 14.3 km NW Goris, above turn c. 1.4 km NNW Tegh , 1275 m, 39 o 34'2''N / 46 o 29'15''E, 23 June 2011, Vitek, Fayvush, Tamanyan, Oganesyan, Margaryan 202284 ( ERE) ; Syunik province, down from vil. Halidzor, 4 September 2019, Nersesyan s.n. ( ERE) .

Section 2. Silvatici Müller (1858: 137).

Stems high- to low-arching, angular, glabrous or pubescent; prickles equal or almost equal, flattened, usually on angles or sometimes displaced; leaves below green or (in our species) grey-tomentose. Inflorescence compound or simple racemose, upward tapering, often with weak glandular pubescence.

Figures

FIGURE 3. Leaves and fragments of stems of Rubus armeniacus (Zangezur, Kafan district, Lower Andes village, at the edge of the field. 9 October 1954. Mulkidzhanyan 66474, ERE)—A, B; R. candicans (Akhtala, Debet river valley. 25 June 1954. Mulkidzhanyan 63240, ERE)—C, D; R. ibericus (Zangezur, outskirts of Goris. 4 September 1949. Karapetyan 135970, ERE)—E, F