Moriomorphina (Eucyclothorax), Liebherr

Liebherr, James K., 2018, Taxonomic review of Australian Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) with special emphasis on the M. lophoides (Chaudoir) species complex, Deutsche Entomologische Zeitschrift 65 (2), pp. 177-224 : 177

Subgenus Eucyclothorax Liebherr

Diagnosis.

Beetles assigned to this subgenus have the prosternum punctate, with punctures either: 1, distributed over the lateral reaches of the prosternum and medial reaches of the proepisternum; 2, more generally distributed across both sclerites; 3, lining a median depression anterad the prosternal process; 4, lining the prosternal-proepisternal suture; or 5, lining a transverse preapical depression ( Liebherr 2018, fig. 2 D–F). The pronotum may be distinctly punctate (Fig. 2 A–B) or not (Fig. 2 C–I), but in all species the median pronotal base is coplanar to only slightly depressed relative to the pronotal disc, not greatly depressed as in Australian species of subgenus Mecyclothorax (Fig. 2 J–M). The vertex is transversely flat to convex, without a transverse dorsal impression, or neck. Generally the species conform to the full complement of standard setae; formula ++/++/+2++. However the basal pronotal seta is absent in M. darlingtoni and M. jameswalkeri , the parascutellar seta is polymorphically present or absent within M. punctatus , and M. isolatus exhibits only a single dorsal elytral seta. Body size ranges from small to moderately large, with standardized body length 2.7-6.0 mm.

All Eucyclothorax spp. are characterized by male genitalia that possess a flagellum, flagellar sheath, and dorsal plate at the apex of the internal sac, as well as robust aedeagal median lobes, i.e. dorsoventrally broad (Figs 7, 14). The female reproductive tract is configured with the spermathecal duct joined ventrally to the juncture of the common oviduct and the bursa copulatrix (Fig. 9). A helminthoid sclerite is present on the bursal wall ventrad the juncture of the spermathecal duct and common oviduct. Like all other Mecyclothorax , the gonocoxae have setae along the apicolateral margin of the basal gonocoxite (Fig. 10), and the apical gonocoxite has 1-3 (usually 2) lateral ensiform setae.